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141. Epigeneium Gagnepain, Bull. Mus. Natl. Hist. Nat., sér. 2. 4: 593. 1932.

厚唇兰属 hou chun lan shu

Authors: Xinqi Chen & Jeffrey J. Wood

Sarcopodium Lindley & Paxton (1850), not Ehrenberg ex Brongniart (1824).

Herbs, epiphytic or terrestrial. Rhizome aerial, creeping and climbing, terminating in a pseudobulb consisting of 1 internode; roots granulose. New shoot arising at base of pseudobulb, sometimes pendulous, covered with conspicuous imbricate chestnut or pale brown sheaths when young. Pseudobulbs usually short and conic, or ovoid, often quadrate, remote or less often contiguous, comprised of 1 internode, usually covered with brown sheaths when young, apex 1- or 2[or 3]-leaved. Leaves distal, shortly petiolate or sessile, oblong, obovate, or elliptic, fleshy, leathery, articulate. Inflorescences pseudoterminal, borne generally on most recent pseudobulb, but occasionally on an unflowered or previously flowered penultimate stem, slender, arising between or just below leaves, 1- to several flowered, usually racemose; floral bracts chestnut, usually much shorter than pedicel and ovary, membranous. Flowers long-lasting, resupinate, medium-sized to large, stellate. Sepals and petals free, narrowly elliptic, subequal. Dorsal sepal enclosing column; lateral sepals larger, oblique and adnate to column foot forming a short, conspicuous mentum. Petals long decurrent on mentum, usually slightly narrower; lip pandurate-oblong, rather narrow, 3-lobed; lateral lobes erect; mid-lobe spreading; disk with a callus lobulate or ridged at base, often of 2 longitudinal lamellae. Column short to rather long, with or without short stelidia, foot long; rostellum subrotund, entire; pollinia 4 in 2 pairs (possibly sometimes only 2), waxy, naked, i.e., without caudicle or viscidium.

About 35 species: Bangladesh, China, Indonesia, Laos, Malaysia, Myanmar, Nepal, New Guinea, Philippines, Thailand, Vietnam; 11 species (four endemic) in China.

The genus Epigeneium was monographed by Seidenfaden (Dansk Bot. Ark. 34(1): 1-104. 1980).

Reviewer L. Averyanov notes that Epigeneium chapaense Gagnepain (Bull. Mus. Natl. Hist. Nat., sér. 2, 4: 596. 1932), described from N Vietnam, also occurs in S China. However, the present authors could not substantiate this record because they found no relevant specimens.


1 Pseudobulbs 1-leaved   (2)
+ Pseudobulbs 2-leaved   (6)
       
2 (1) Lip rounded and apiculate or slightly emarginate at apex   (3)
+ Lip conspicuously bilobed at apex   (4)
       
3 (2) Lip with 2 keels in lower half united apically into a small transverse callus with 3 low keels in front and ± continuing to apex.   4 E. nakaharae
+ Lip bearing only 3 low keels restricted to middle.   5 E. mimicum
       
4 (2) Calli convergent; lateral sepals ca. 9 mm wide; lip epichile ca. 10 mm wide, distinctly wider than hypochile; leaves 2.8-4.6 cm.   3 E. tsangianum
+ Calli separate; lateral sepals 5-6 mm wide; lip and leaves without above combination of characters   (5)
       
5 (4) Leaves 1-2.3 cm; lip epichile ca. 11 mm wide, apex deeply bilobed; sepals and petals pale pink or reddish orange, lip white.   1 E. fargesii
+ Leaves 5-6 cm; lip epichile 8-9 mm wide, apex shallowly bilobed or notched; flowers dull purple to brown.   2 E. clemensiae
       
6 (1) Dorsal sepal 45-65 mm; leaves (6-)10-22.5 cm.   6 E. amplum
+ Dorsal sepal 15-32 mm; leaves 2.5-9.5 cm   (7)
       
7 (6) Petals 5-6 mm wide; sepals 22-32 mm; mid-lobe of lip subreniform to obscurely bilobed, emarginate.   7 E. rotundatum
+ Petals less than 4 mm wide; sepals 15-25 mm; mid-lobe not as above   (8)
       
8 (7) Disk with 3 indistinct keels and a distal callus.   8 E. gaoligongense
+ Disk with 3 prominent fleshy carinate lamellae, distal callus absent   (9)
       
9 (8) Lateral lobes of lip broad; mid-lobe of lip minutely mucronate to hooked aristate at apex.   9 E. fuscescens
+ Lateral lobes of lip narrow; mid-lobe of lip rounded at apex   (10)
       
10 (9) Lateral keels on disk most prominent, obliquely truncate apically, median keel very low.   10 E. treutleri
+ All keels similar in size and shape in upper half, before fusing in apical quarter, each one terminating in a 2-angled manner.   11 E. forrestii

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