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FOC | Family List | FOC Vol. 24 | Eriocaulaceae

1. Eriocaulon Linnaeus, Sp. Pl. 1: 87. 1753; Gen. Pl. ed. 5; 38, 1754.

谷精草属 gu jing cao shu

Herbs scapigerous, aquatic or growing in marshes, rarely on dry ground. Stems very short and disciform, rarely elongate. Leaves radical, linear. Scapes 3--8-ribbed; sheath oblique at mouth. Bracts scalelike. Flowers unisexual, with both sexes in same head, minute. Male flowers: sepals 2 or 3, usually connate into an abaxially split tube, rarely free; petals basally connate and funnelform, large, margin usually ciliate, apex distinctly or indistinctly 2- or 3-lobed, usually bearing a gland; stamens in 2 whorls, adnate to petals; anthers black or rarely yellowish. Female flowers: sepals 2 or 3, free, basally connate, or connate into an adaxial spathe, flat, boatlike, or crested, equal or unequal; petals absent to 3, linear, oblanceolate, or spatulate, margin ciliate, apex often bearing a large gland; ovary 1--3-loculed; style 1--3-branched. Seeds yellowish or brown, 1 per valve; testa usually hexagonally reticulate, prickles small and in rows or absent; embryo minute.

About 400 species: mainly in tropical and subtropical regions, with a concentration in Asia; 35 species (13 endemic) in China.

Eriocaulon occurs throughout most of China except for Ningxia, Qinghai, and Xinjiang.


1 Flowers 2-merous; ovary 2-loculed but sometimes only 1 locule developed   35 ERIOCAULON decemflorum
+ Flowers 3-merous but sometimes parts fused or degenerate; ovary 3-loculed or 1 or 2 locules not developed.   (2)
       
2 (1) Sepals of female flowers connate at base or spathelike, apex 3-lobed.   (3)
+ Sepals of female flowers free (sometimes ± connate in E. tonkinense).   (11)
       
3 (2) Ovary 1(or 2)-loculed.   (4)
+ Ovary 3-loculed.   (5)
       
4 (3) Scapes 20--35 cm; leaves 4--12 cm; sheath 3--7 cm   33 ERIOCAULON faberi
+ Scapes ca. 3.5 cm; leaves 1--2.8 cm; sheath 0.6--1.7 cm   34 ERIOCAULON mangshanense
       
5 (3) Floral bract abaxially white clavate hairy toward apex.   (6)
+ Floral bract glabrous, glabrescent, margin villous, or abaxially shortly hairy toward apex.   (7)
       
6 (5) Receptacle usually densely villous; head straw-colored, 3--5 × 4--5 mm; involucral bracts orbicular to obovate, 2--2.5 mm; seed prickles T-shaped   26 ERIOCAULON buergerianum
+ Receptacle glabrous or with very few short hairs; head yellowish green, ca. 2.5 × 4 mm; involucral bracts broadly ovate, 1.5--2 mm; seed prickles linear, apex truncate or 2-branched   30 ERIOCAULON chinorossicum
       
7 (5) Leaves 0.5--0.7 mm wide at middle   32 ERIOCAULON minusculum
+ Leaves 1--5 mm wide at middle.   (8)
       
8 (7) Petals of female flowers glabrous   31 ERIOCAULON glabripetalum
+ Petals of female flowers adaxially villous.   (9)
       
9 (8) Leaves 1--1.5 mm wide at middle, 4--7-veined; involucral bracts lanceolate to linear-lanceolate, 3.5--7.3 mm   27 ERIOCAULON miquelianum
+ Leaves 2--5 mm wide at middle, 7--20-veined; involucral bracts ovate to oblong, 1.5--2 mm.   (10)
       
10 (9) Leaf veins 12--20; heads turbinate; seed prickles 1 or 2(--4) per cell   28 ERIOCAULON alpestre
+ Leaf veins 7--12; heads subglobose; seed prickles 2--4 per cell   29 ERIOCAULON robustius
       
11 (2) Petals of female flowers absent.   (12)
+ Petals of female flowers present.   (14)
       
12 (11) Apex of involucral bracts acuminate to caudate; sepals of female flowers winged   22 ERIOCAULON echinulatum
+ Apex of involucral bracts obtuse to acute; sepals of female flowers wingless.   (13)
       
13 (12) Sepals of male flowers spathelike, apex 3-cleft; anthers white to milky white or dusky; leaves ca. 1 mm wide at middle   20 ERIOCAULON cinereum
+ Sepals of male flowers tubular at base, apex 2-cleft; anthers blackish; leaves 1.8--2 mm wide at middle   21 ERIOCAULON taishanense
       
14 (11) Sepals of female flowers usually 2, sometimes 3 on flowers near edge of inflorescence.   (15)
+ Sepals of female flowers 3 (rarely 2 in E. sexangulare).   (16)
       
15 (14) Leaves 6--8 cm, subleathery; scapes 1 or 2, 7-ribbed   19 ERIOCAULON sclerophyllum
+ Leaves 2--5 cm, herbaceous or membranous and conspicuously fenestrate; scapes 5--10, (4 or)5(or 6)-ribbed   18 ERIOCAULON truncatum
       
16 (14) Stem elongate.   (17)
+ Stem not elongate.   (18)
       
17 (16) Leaves ca. 0.2 mm wide at middle, 1-veined; sheath 2--2.5 cm; heads ca. 3 mm in diam.; involucral bract ca. 1 mm   10 ERIOCAULON setaceum
+ Leaves 3--5 mm wide at middle, 12--28-veined; sheath 3--6 cm; heads ca. 5 mm in diam.; involucral bract 2--2.7 mm   12 ERIOCAULON nepalense
       
18 (16) Floral bracts abaxially white clavate hairy toward apex.   (19)
+ Floral bracts glabrous, ciliate, or abaxial hairs not white and clavate.   (27)
       
19 (18) Involucral bracts villous at least at base abaxially.   (20)
+ Involucral bracts glabrous or subglabrous.   (21)
       
20 (19) Leaves 4--6 mm wide; involucral bracts oblong, 3.5--4 × 2.2--3.6 mm; lateral sepals wingless   6 ERIOCAULON brownianum
+ Leaves 2--3 mm wide; involucral bracts orbicular-reniform to ovate, 2--2.5 × 2--2.5 mm; lateral sepals winged   7 ERIOCAULON australe
       
21 (19) Involucral bracts brownish, suborbicular   3 ERIOCAULON angustulum
+ Involucral bracts straw-colored (sometimes blackish at apex), elliptic, ovate, or obovate.   (22)
       
22 (21) Gland of male flower petals absent, inconspicuous, or present only on lateral petals.   (23)
+ Gland of male flower petals present.   (24)
       
23 (22) Leaves 3--3.5 mm wide at middle; sheath 5.5--8 cm   1 ERIOCAULON acutibracteatum
+ Leaves 1--1.8(--3) mm wide at middle; sheath 2.5--4 cm   2 ERIOCAULON nantoense
       
24 (22) Heads white, usually with black, triangular patches, 5--6 mm in diam.   9 ERIOCAULON schochianum
+ Heads black or blackish, 3--5 mm in diam.   (25)
       
25 (24) Anthers milky white to brownish; seed prickles absent   14 ERIOCAULON tonkinense
+ Anthers black; seed prickles linear.   (26)
       
26 (25) Leaves 1--1.8(--3) mm wide at middle; heads 3.5--5 mm in diam.   2 ERIOCAULON nantoense
+ Leaves ca. 3 mm wide at middle; heads 3--3.5 mm in diam.   11 ERIOCAULON sollyanum
       
27 (18) Receptacle glabrous or subglabrous.   (28)
+ Receptacle sparsely to densely hairy.   (30)
       
28 (27) Sepals of male flowers laterally winged; seed prickles T-shaped   8 ERIOCAULON sexangulare
+ Sepals of male flowers wingless; seed prickles absent.   (29)
       
29 (28) Sepals of female flowers keeled; petals of female flowers glabrous, apex emarginate   15 ERIOCAULON kunmingense
+ Sepals of female flowers not keeled; petals of female flowers adaxially villous at center, apex acute   16 ERIOCAULON leianthum
       
30 (27) Male and female flower petals with a gland (sometimes gland on only 2 petals of female flowers in E. henryanum).   (31)
+ Male and female flower petals without a gland (occasionally gland present on lateral petals of female flowers in E. rockianum).   (32)
       
31 (30) Leaves 4--8 cm; scapes 1--4; sheath 4--9 cm   4 ERIOCAULON henryanum
+ Leaves ca. 2.3 cm; scapes 5--23; sheath 2--2.3 cm   23 ERIOCAULON parvum
       
32 (30) Floral bracts glabrous, shortly hairy, or margin ciliate.   (33)
+ Floral bracts villous, hoary, or hispid.   (34)
       
33 (32) Involucral bracts 2.2--2.5 mm; male flower petals usually glabrous   17 ERIOCAULON zollingerianum
+ Involucral bracts ca. 2 mm; male flower petals hoary or white clavate hairy   25 ERIOCAULON ermeiense
       
34 (32) Sheath 5.5--10 cm; involucral bracts 3--4 × 2.5--3 cm; floral bracts hispid; seed prickles setiform, 2 or 3 from transverse wall and 1 from longitudinal wall   24 ERIOCAULON oryzetorum
+ Sheath 1--5 cm; involucral bracts 2--3 × 1.3--2 cm; floral bracts villous or hoary; seed prickles oblong or linear.   (35)
       
35 (34) Sheath 1--2.5 cm; heads ca. 2 mm; involucral bracts ca. 2.2 mm, membranous; floral bracts villous; prickles oblong   5 ERIOCAULON rockianum
+ Sheath 2.5--5 cm; heads 3.5--5 mm; involucral bracts 2.5--3 mm, papery; floral bracts hoary; prickles linear   13 ERIOCAULON luzulifolium

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