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56. Faberia Hemsley, J. Linn. Soc., Bot. 23: 479. 1888.

花佩菊属 hua pei ju shu

Authors: Zhu Shi & Norbert Kilian

Faberiopsis C. Shih & Y. L. Chen.

Herbs, perennial, often rosulate, with rhizomes. Stem leafy or ± leafless. Leaves lyrately pinnate or undivided, leathery. Capitula with 5-30 florets. Involucre ± narrowly cylindric to ± narrowly campanulate. Phyllaries mostly glabrous; outer phyllaries in several series, gradually longer centripetally, often conspicuously imbricate, longest ca. 1/2 as long as inner ones; inner phyllaries 5-14, ± equal in length, ± linear-lanceolate to linear. Receptacle naked. Florets reddish to bluish purple. Achene brown to reddish brown, subcylindric to narrowly ellipsoid, rather weakly compressed, with 5 main ribs and 1 or 2 narrower ribs in between, apex truncate. Pappus brownish, single, of strong scabrid bristles.

● Seven species: China.

Systematic placement and circumscription of the genus have been revised based on molecular phylogenetic analyses of subtribes Lactucinae and Crepidinae (J. W. Zhang & N. Kilian, in prep.; N. Kilian et al., in prep.). N. Kilian et al. (in V. A. Funk et al., Syst. Evol. Biogeogr. Compositae, 348-350. 2009) placed Faberia in subtribe Crepidinae, but now ITS phylogenies show it on a very basal branch in subtribe Lactucinae. Recent additions to the genus by Sennikov (Komarovia 5: 109-110. 2008) are based on a different genus concept and are, as far as relevant for the flora of China, not supported. Karyological studies (Y. Liu, T. Deng & Q. E. Yang, pers. comm.) have revealed that the four species investigated have the unusual basic chromosome number of x = 17, perhaps indicating a hybrid origin of this genus.

Prenanthes glandulosa Dunn (J. Linn. Soc., Bot. 35: 514. 1903), which was initially placed by the first author in Notoseris (N. glandulosa (Dunn) C. Shih), is only known from the holotype at K, a piece of a flowering plant raised from seeds collected in "West China." It seems to be a distinct species, perhaps referable to Faberia, as may be assumed from the involucre (purplish, inner phyllaries ca. 6, outer ones linear as in F. lancifolia), the 10-12 blue (or purple?) florets, and the pale straw-colored pappus. The lower leaves have a broadly ovate blade with cordate base and a distinct, basally sheathlike widened and clasping petiole. The axes of the paniculiform to corymbiform synflorescence are densely glandular hairy, a feature otherwise not known from Faberia.

1 Leaves lyrately pinnatifid to lyrately pinnatisect   (2)
+ Leaf blade not divided or at most coarsely sinuate-dentate   (3)
2 (1) Leaves with terminal lobe much larger than lateral lobes, to 2/3 of entire leaf; involucre 1.4-1.5 cm; inner phyllaries usually 12-14; capitula with 20-30 florets.   3 F. sinensis
+ Leaves with terminal lobe never conspicuously larger than lateral lobes; involucre 1.1-1.3 cm; inner phyllaries ca. 8; capitula with 10-15 florets.   4 F. ceterach
3 (1) Leaf blade lanceolate to narrowly elliptic or oblanceolate, at least ca. 3 × as long as wide   (4)
+ Leaf blade broadly elliptic, ovate, or pentagonal, mostly not more than ca. 2 × as long as wide   (5)
4 (3) Capitula many in a leafless narrowly paniculiform synflorescence.   1 F. lancifolia
+ Capitula 1-3 on short branchlets subtended by upper stem leaves.   2 F. nanchuanensis
5 (3) Leaves on stem, with petiole usually shorter than blade; leaf blade triangular-ovate to pentagonal; capitula with 5 florets; inner phyllaries 5.   7 F. faberi
+ Most leaves basal, with petiole distinctly longer than blade; leaf blade broadly elliptic to ovate; capitula with 10-20 florets; inner phyllaries at least 8   (6)
6 (5) Leaf blade mostly more than 5 cm; synflorescence corymbiform, with some capitula; florets bluish purple.   5 F. cavaleriei
+ Leaf blade at most ca. 4 cm; synflorescence sparsely branched, with 1-4 capitula only; florets purplish, drying yellowish.   6 F. thibetica

Lower Taxa


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