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FOC | Family List | FOC Vol. 18 | Scrophulariaceae

10. Gratiola Linnaeus, Sp. Pl. 1: 17. 1753.

水八角属 shui ba jiao shu

Herbs, perennial or annual, erect or prostrate, succulent, glabrous or glandular hairy. Leaves opposite, sessile, margin entire or serrate. Flowers axillary, solitary. Pedicel short or long, filiform. Bracteoles 2, foliaceous, near calyx. Calyx deeply 5-lobed; lobes narrow, subequal, imbricate. Corolla 2-lipped to nearly so, lips spreading; lower lip 3-lobed; upper lip entire to 2-lobed. Stamens 2, inserted on upper side of corolla tube, included; anther locules distinct, parallel, transverse or erect; staminodes 2 and included, or absent. Style filamentous; stigma 2-lamellate, included. Capsule ovoid to globose, septifragal, 4-valved; valves separating from placental axis, apex acute. Seeds numerous; seed coat striate and transversely reticulate.

About 25 species: mostly temperate and tropics; three species in China.

1 Leaf blade apex obtuse; staminodes absent; capsule ovoid, 2.5-3 mm in diam.   2 Gratiola griffithii
+ Leaf blade apex acute to apiculate; staminodes 2; capsule globose, 4-5 mm in diam.   (2)
2 (1) Flowers pedicellate, pedicel 0.7-1.5 cm; corolla 1-1.3 cm   3 Gratiola officinalis
+ Flowers sessile or subsessile; corolla 0.5-0.7 cm   1 Gratiola japonica

Lower Taxa


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