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  | 1 | Roots dilated into tuberiform rhizomes |  | (2) |  
  | + | Roots not dilated, not tuberiform |  | (3) |  
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  | 2 (1) | Scapigerous herbs; flowering stem simple, 20-50 cm tall; leaves crowded at base, rosulate, obovate, spatulate, or elliptic, marginally repand-dentate or pinnatifid; petioles basally not auriculate; capitula 1-5, in lax corymb. |  | 1 G. pseudochina |  
  | + | Robust herbs, to 150 cm tall, much branched; stem leaves numerous, lyrate-pinnatipartite to pinnatipartite; petioles basally rounded, with pinnatifid auricles; capitula numerous, corymbose-paniculate. |  | 2 G. japonica |  
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  | 3 (1) | Scandent herbs; stems procumbent; leaves ovate or ovate-oblong, apically acute or acuminate, marginally entire or repand-dentate, both surfaces glabrous or sparsely hairy, base of petioles not auriculate; capitula 3-5 in terminal or axillary corymbs. |  | 10 G. procumbens |  
  | + | Erect herbs or shrubs |  | (4) |  
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  | 4 (3) | Stem procumbent in lower part |  | (5) |  
  | + | Stem erect or declined at base |  | (7) |  
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  | 5 (4) | Plants and phyllaries densely fulvous tomentose; leaves narrowly elliptic, ovate, or rhombic, margin entire or serrate in upper half, basally cuneate-attenuate into petiole; lateral veins 3-6-paired, both surfaces sparsely or densely shortly hairy. |  | 3 G. nepalensis |  
  | + | Plants and phyllaries glabrous or subglabrous |  | (6) |  
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  | 6 (5) | Leaves sessile or subsessile, oblong-elliptic, obovate, or oblong-lanceolate, basally amplexicaul, with broad auricle, margin irregularly sharply serrate, lateral veins 12-30-paired; capitula 4-15, corymbose-paniculate. |  | 5 G. cusimbua |  
  | + | Leaves petiolate, obovate or oblanceolate, basally cuneate, narrow, not auriculate, margin repand-dentate, or mucronulate, lateral veins 7-9-paired; capitula numerous, laxly corymbose. |  | 4 G. bicolor |  
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  | 7 (4) | Leaves simple, elliptic, marginally entire or laxly denticulate, apex obtuse, base of petiole with incised auricle. |  | 9 G. elliptica |  
  | + | Leaves marginally repand-dentate or lyrately pinnately divided |  | (8) |  
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  | 8 (7) | Leaves usually crowded on lower part of stem, petiolate or subsessile, ovate, elliptic, or oblanceolate, veinlets coalescent into suboblong, parallel-reticulate, conspicuous black lines when dry; capitula 3-5 corymbose-paniculate, divaricate. |  | 6 G. divaricata |  
  | + | Leaves lax, rarely crowded on stem, elliptic or spatulate, veinlets inconspicuous, not coalescent into black lines when dry; capitula laxly corymbose, not divaricate |  | (9) |  
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  | 9 (8) | Plants hispid; leaves lyrate-ovate, basally attenuate into long petioles, upper or median leaves 1- or 2-denticulate, apically obtuse, both surfaces adpressed pilose; capitula 3, laxly corymbose; peduncles 5-7 cm. |  | 7 G. formosana |  
  | + | Plants fulvous pubescent; leaves lyrate-pinnatilobate, 4-12 cm, terminal lobe large, triangular-ovate, apex acuminate, base truncate or cordate, lateral lobes 1- or 2-paired, ovate-oblong, both surfaces fulvous pubescent; capitula 1-3, laxly corymbose; peduncles 1-3 cm. |  | 8 G. barbareifolia |  |