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19. Haplopteris C. Presl, Tent. Pterid.  141.1836.  
书带蕨属 shu dai jue shu 
 
Authors: Zhang Xianchun & Michael G. Gilbert
 
 
 
 
Epiphytic or epilithic grasslike plants. Fronds simple, linear, entire, glabrous. Venation consisting of a costa and oblique unbranched veins, free except for a submarginal fertile connecting vein. Sori in one elongate marginal or submarginal groove on each side of lamina, immersed, rarely superficial; paraphyses long with a dark obconic head. Spores monolete, most ellipsoid, sometimes fusiform, smooth, transparent. 
About 40 unclear species: tropical and subtropical Asia; 13 species in China. 
 
 
 
Most of the Old World species formally classified in Vittaria now belong to Haplopteris. Vittaria suberecta Hayata (Icon. Pl. Formosan. 6: 161. 1916), described from Taiwan, could not be treated here because no material was seen by the present authors. 
 
 
 
 
                        
                         
		
                          
                          
			
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  | 1 | 
  Soral line marginal, immersed in groove, open outward | 
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  (2) | 
 
  | + | 
  Soral line submarginal | 
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  (3) | 
 
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  | 2 (1) | 
  Lamina up to 100 cm or longer, ca. 10 mm wide; stipe long, slender; scales dark brown. | 
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  12 H. elongata | 
 
  | + | 
  Lamina 8-30 cm, 2-4 mm wide; stipe short; scales yellow-brown. | 
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  13 H. anguste-elongata | 
 
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  | 3 (1) | 
  Soral line superficial; lamina margins straight or revolute | 
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  (4) | 
 
  | + | 
  Soral line immersed in groove between costa and margins of lamina; lamina margins revolute | 
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  (8) | 
 
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  | 4 (3) | 
  Scales dark brown, small, straight, areole wall thick; soral line submarginal, covered by revolute margins of lamina. | 
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  1 H. amboinensis | 
 
  | + | 
  Scales yellow-brown, large, soft, twisted, areole wall thin; soral line with a broad area between margins of lamina | 
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  (5) | 
 
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  | 5 (4) | 
  Lamina 2-3 mm wide; scales 3-7 ?ca. 0.2 mm; soral line close to costa. | 
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  5 H. mediosora | 
 
  | + | 
  Lamina over 5 mm wide; scales 10-20 ?0.5-2.5 mm; soral line with a broad area between costa | 
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  (6) | 
 
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  | 6 (5) | 
  Stipe short, thick; lamina 10-30 mm wide; scales 10-20 mm. | 
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  2 H. doniana | 
 
  | + | 
  Stipe absent, or slender; lamina 5-12 mm wide; scales ca. 10 mm | 
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  (7) | 
 
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  | 7 (6) | 
  Lamina 6-12 mm wide; soral line 1-1.5 mm away from margins of lamina. | 
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  3 H. taeniophylla | 
 
  | + | 
  Lamina 5-6 mm wide; soral line ca. 1 mm away from margins of lamina. | 
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  4 H. himalayensis | 
 
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  | 8 (3) | 
  Lamina 5-10 mm wide, or more | 
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  (9) | 
 
  | + | 
  Lamina (0.5-)2-4(-11) mm wide | 
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  (10) | 
 
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  | 9 (8) | 
  Costa not raised adaxially, lamina with a narrow concave line adaxially. | 
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  6 H. plurisulcata | 
 
  | + | 
  Costa raised adaxially, lamina with a broad concave line on each side of costa adaxially. | 
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  7 H. fudzinoi | 
 
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  | 10 (8) | 
  Scales dark or light brown, 5-10 mm | 
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  (11) | 
 
  | + | 
  Scales yellow-brown, 2-3 mm | 
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  (12) | 
 
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  | 11 (10) | 
  Scales light brown; lamina with soral line very close to costa. | 
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  8 H. linearifolia | 
 
  | + | 
  Scales dark brown; lamina with soral line close to margins. | 
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  9 H. hainanensis | 
 
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  | 12 (10) | 
  Costa flattened abaxially, wide; soral line covered by costa. | 
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  10 H. sikkimensis | 
 
  | + | 
  Costa raised abaxially, narrow; soral line with a broad area between costa. | 
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  11 H. flexuosa | 
 
 
 
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                          List of lower taxa 
 
                        
                          
                        
                        
                      
 
                      
	 
                      
                         
		
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