All Floras      Advanced Search
FOC Vol. 5 Page 192 Login | eFloras Home | Help
FOC | Family List | FOC Vol. 5 | Proteaceae

1. Helicia Loureiro, Fl. Cochinch. 1: 83. 1790.

山龙眼属 shan long yan shu

Trees, rarely shrubs. Leaves alternate, rarely ± opposite or ± whorled, petiolate or subsessile, simple, margin entire or serrate. Inflorescence axillary, ramiflorous, rarely subterminal, a raceme of flower pairs, many-flowered. Flowers bisexual; bracts of flowers and flower pairs small, ovate-acuminate to subulate, rarely foliaceous, caducous, rarely persistent. Flower pairs sessile (“pedicels free”) or with a basal common peduncle (“pedicels connate”). Perianth tube straight, slender, limb clavate to ± globose, segments free and revolute at anthesis. Stamens subsessile, attached slightly below limb; anthers oblong; connective apiculate. Pollen grains triporate, exine scabrate-granulate. Hypogynous glands 4, free or connate into a ring or cup. Ovary sessile; ovules 2, anatropous, inserted at base of locule or laterally on wall. Style slender; stigma terminal, punctiform. Fruit usually indehiscent, although sometimes splitting along ventral suture; pericarp mostly thick and leathery. Seeds 1 or 2, ± globose to hemispheric; testa thin; cotyledons fleshy, apically rugulose.

About 97 species: from S India and Japan to SE Australia, mainly in SE Asia and New Guinea; 20 species (ten endemic) in China.


1 Ovary densely hairy.   (2)
+ Ovary glabrous.   (3)
       
2 (1) Young leaves and flowers flexuose rust-colored tomentose; fruit brown, ± globose.   1 H. vestita
+ Young leaves and flowers rust-colored tomentulose; fruit purplish black, obovoid to ellipsoid.   2 H. obovatifolia
       
3 (1) Branchlets (apices excepted in some species) and leaves glabrous when young.   (4)
+ Branchlets and leaves densely hairy when young.   (11)
       
4 (3) Branchlets apically glabrous.   (5)
+ Branchlets apically hairy.   (7)
       
5 (4) Fruit apex not beaked; pericarp leathery, ca. 0.5 mm thick.   17 H. cochinchinensis
+ Fruit apex markedly beaked; pericarp woody, 1-1.5 mm thick.   (6)
       
6 (5) Leaf blade leathery, margin entire or remotely denticulate; petiole 10-20 mm; rachis and pedicel pilosulose; pedicel 2.5-3 mm; perianth ca. 2.5 cm, pilosulose.   11 H. dongxingensis
+ Leaf blade papery, margin serrate on apical half; petiole 1-6 mm; rachis and pedicel glabrous; pedicel 3-5 mm; perianth 1.5-2.2 cm, glabrous.   12 H. hainanensis
       
7 (4) Bracts of flower pairs 4-10 mm.   18 H. shweliensis
+ Bracts of flower pairs 1-3 mm.   (8)
       
8 (7) Leaf blade with reticulate veins conspicuous on both surfaces.   (9)
+ Leaf blade with reticulate veins conspicuous only on adaxial surface.   (10)
       
9 (8) Perianth 1.3-1.6 cm; hypogynous glands basally connate.   14 H. reticulata
+ Perianth 1-1.4 cm; hypogynous glands free.   15 H. rengetiensis
       
10 (8) Petiole 2-4.5 cm; perianth white, 1.6-2.5 cm; hypogynous glands connate.   19 H. longipetiolata
+ Petiole 0.8-2 cm; perianth greenish white or reddish, 1.2-1.6 cm; hypogynous glands free.   20 H. falcata
       
11 (3) Leaves and rachis of inflorescences hairy when young, mature leaf blade glabrescent.   (12)
+ Leaves and rachis of inflorescence rust-colored tomentose when young, mature leaf blade adaxially hairy or with hairs along midvein.   (16)
       
12 (11) Reticulate veins inconspicuous.   16 H. kwangtungensis
+ Reticulate veins conspicuous.   (13)
       
13 (12) Pericarp woody.   (14)
+ Pericarp leathery.   (15)
       
14 (13) Bracts of flower pairs ovate, ca. 1 mm; pedicel glabrous, 3-4 mm; perianth 1.2-1.5 cm.   9 H. tsaii
+ Bracts of flower pairs lanceolate, ca. 2 mm; pedicel pilosulose, 6-7 mm; perianth 1.6-1.8 cm.   10 H. tibetensis
       
15 (13) Bracts of flower pairs ca. 1 mm, pilose; fruit dark green, ± globose, 2.5-3.5 cm in diam.; pericarp 1.5-3 mm thick.   13 H. nilagirica
+ Bracts of flower pairs 1.5-2 mm, glabrous; fruit black, broadly ellipsoid, ca. 1.5 cm in diam.; pericarp ca. 1 mm thick.   14 H. reticulata
       
16 (11) Perianth 2-3 cm; rachis 2-4 mm in diam. at anthesis.   (17)
+ Perianth 1-2 cm; rachis 1-2.5 mm in diam. at anthesis.   (18)
       
17 (16) Petiole 0.2-1.5 cm; leaf blade ± leathery, abaxially tomentose.   3 H. grandis
+ Petiole 2.5-3 cm; leaf blade papery, abaxially pilose along midvein.   4 H. pyrrhobotrya
       
18 (16) Perianth 1-1.4 cm, glabrous; pericarp leathery.   (19)
+ Perianth 1.4-2 cm, pilose; pericarp woody.   (20)
       
19 (18) Mature leaf blade oblanceolate, adaxially pilose along midvein, margin dentiform serrate; rachis 1.5 mm in diam.   5 H. silvicola
+ Mature leaf blade broadly obovate-oblong to obovate, both surfaces pilose along midvein, margin remotely serrate or rarely entire; rachis 2.5 mm in diam.   6 H. clivicola
       
20 (18) Perianth 1.5-2 cm; mature leaf blade abaxially pilose along all veins, margin toothed; inflorescences 10-24 cm.   7 H. formosana
+ Perianth 1.4-1.5 cm; mature leaf blade abaxially pilose along midvein, margin entire; inflorescences 7-8 cm.   8 H. yangchunensis

  • List of lower taxa


     

    Related Objects  
  • PDF File
  • PDF

  • Flora of China @ efloras.org
    Browse by
    Volume
    Family
    Genera
    Advanced Search


    Flora of China Home


    Checklist

     

     

     |  eFlora Home |  People Search  |  Help  |  ActKey  |  Hu Cards  |  Glossary  |