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FOC | Family List | FOC Vol. 14 | Helwingiaceae

1. Helwingia Willdenow, Sp. Pl. 4: 634, 716. 1806, nom. cons., not Helvingia Adanson (1763).

青荚叶属 qing jia ye shu

Helwingia japonica

Credit: Harvard University Herbaria

Shrubs evergreen or deciduous, 1–2 m tall, rarely small trees to 8 m tall. Leaves petiolate; petiole rounded; leaf blade linear-lanceolate to broadly ovate, papery, subleathery, or leathery, glabrous or pubescent; veins 5–9, inconspicuous or conspicuous. Flowers ca. 3–5 mm in diam. Petals green or reddish purple, triangular ovate, valvate. Fruit green when young, turning red and eventually black when mature, subglobose, ovoid, or oblong, green when young.

Geographical distribution is the same as that of the family.

Helwingia argyi H. Léveillé & Vaniot is a synonym of Stemona japonica (Blume) Miquel in the Stemonaceae (see Fl. China 24: 70. 2000).

1 Shrubs, rarely small trees; leaves papery or thickly papery; veins on adaxial surface conspicuous.   (2)
+ Small trees or shrubs; leaves leathery or subleathery, rarely thickly papery; veins on adaxial surface inconspicuous.   (3)
2 (1) Leaves ovate, ovate-rounded, or broadly elliptic, rarely ovate lanceolate; stipules linear or filiform, divided, rarely undivided; secondary and smaller lateral veins of leaves conspicuous.   1 Helwingia japonica
+ Leaves narrowly elliptic or oblong lanceolate, rarely ± diamond-shaped; ratio of length to width 3 or greater; stipules often 2(or 3)-divided, rarely undivided; secondary and smaller lateral veins of leaves inconspicuous.   2 Helwingia himalaica
3 (1) Leaves linear-lanceolate or lanceolate, 4–15 × 0.4–2 cm, margins sparsely glandular serrate.   3 Helwingia chinensis
+ Leaves obovate-oblong, oblong, rarely oblanceolate, larger, 9–15 × 3–5 cm, margins densely glandular serrate.   4 Helwingia omeiensis

Lower Taxa


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