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FOC | Family List | FOC Vol. 18 | Gesneriaceae

25. Hemiboea C. B. Clarke, Hooker's Icon. Pl. 18: sub pl. 1798.

半蒴苣苔属 ban shuo ju tai shu

Herbs, perennial, terrestrial or epipetric, stoloniferous. Stems simple or branched. Leaves few to many, along stem, opposite, equal to subequal in a pair; leaf blade glabrous to pilose, base attenuate to cuneate, rarely rounded. Inflorescences umbel-like, dense, axillary or pseudoterminal, 1- to many-flowered cymes; bracts 2, opposite, connate, forming a globose to ovoid involucre. Calyx actinomorphic, 5-sect from base, 2-lipped, or 5-lobed from middle or above; segments equal, rarely unequal. Corolla white to pink, purple, or yellow, zygomorphic, inside usually with a ring of hairs; tube funnelform-tubular, not swollen, longer than limb, 0.7-2 cm in diam.; limb 2-lipped; adaxial lip 2-lobed, shorter than to nearly as long as abaxial lip; abaxial lip 3-lobed, lobes equal or subequal, apex rounded. Stamens 2, adnate to abaxial side of corolla tube below middle, included; anthers basifixed, coherent apically or adaxially, thecae parallel, not confluent at apex, dehiscing longitudinally; connective not projecting; staminodes 2 or 3, adnate to adaxial side of corolla tube. Disc ringlike. Ovary linear to linear-lanceolate, 2-loculed, only adaxial locule fertile; placenta 1, axile. Stigma 1, terminal, truncate to obtuse or capitate, undivided. Capsule straight or oblique in relation to pedicel, usually narrowly lanceolate, somewhat curved, much longer than calyx, dehiscing loculicidally to base only adaxially; valves 2, straight, not twisted. Seeds unappendaged.

Twenty-three species: China, S Japan, N Vietnam; all 23 species in China.

Petioles are often described as winged in Hemiboea; since a winged petiole is really a decurrent leaf blade, they are described here as the leaf blade base.

1 Calyx 5-lobed from middle or above, lobes equal.   (2)
+ Calyx 5-sect from base, segments equal, or unequal and 2-lipped with adaxial lip 2-lobed from base or below middle.   (5)
2 (1) Calyx 5-lobed from middle; corolla pink outside   20 Hemiboea gamosepala
+ Calyx 5-lobed from above middle; corolla white outside.   (3)
3 (2) Leaf blade without sclereids; petiole 1-10.5 cm; peduncle 1-4.5 cm; corolla 1.5-1.8 cm, glabrous outside; staminodes 1-2.5 mm; pistil ca. 0.9 cm   22 Hemiboea follicularis
+ Leaf blade with sclereids; petiole 0.5-4 cm; peduncle 0.6-1.5 cm; corolla 3.7-4.5 cm, sparsely glandular puberulent outside; staminodes ca. 8 mm; pistil ca. 2.6 cm.   (4)
4 (3) Stems much branched, with 7 to many nodes; leaf blade 3-10 X 1-4 cm, apex cuspidate to acuminate; involucre ca. 1 cm in diam.; calyx lobes 3-4 X 2-3 mm; anthers ca. 2.5 mm   21 Hemiboea wangiana
+ Stems unbranched, with 3-5 nodes; leaf blade 8-24 X 4-11 cm, apex acute; involucre 2.5-3 cm in diam.; calyx lobes 7-10 X 8-10 mm; anthers ca. 4 mm   23 Hemiboea magnibracteata
5 (1) Calyx segments outside and margin puberulent or glandular puberulent.   (6)
+ Calyx segments outside and margin glabrous.   (13)
6 (5) Ovary glandular puberulent or white pilose.   (7)
+ Ovary glabrous.   (9)
7 (6) Stems sparsely spreading glandular pubescent toward apex, with 10-15 nodes; leaf blade abaxially glabrous; lateral veins 8-12 on each side of midrib; peduncle 1-2 cm; corolla pale yellow, with a ring of hairs inside   19 Hemiboea longgangensis
+ Stems densely puberulent to villous, with 3-7 nodes; leaf blade abaxially pubescent to densely pubescent; lateral veins 3-8 on each side of midrib; peduncle 0.4-5.5(-10) cm; corolla white, glabrous inside.   (8)
8 (7) Cymes 2-7(-21)-flowered; involucre 1-2.5 cm in diam.; anthers ca. 3.2 mm; staminodes 2, 6-8 mm   17 Hemiboea flaccida
+ Cymes 2- or 3-flowered; involucre 0.4-0.5 cm in diam.; anthers ca. 1.5 mm; staminodes 3, 0.4-2.5 mm   18 Hemiboea parvibracteata
9 (6) Leaf blade apex obtuse; staminodes 3, 0.6-2.8 mm; cymes 3-10- or more flowered; adaxial corolla lip ca. 3.5 mm   16 Hemiboea strigosa
+ Leaf blade apex acute to acuminate; staminodes 2, 4-7 mm; cymes 1-6-flowered; adaxial corolla lip 5-10 mm.   (10)
10 (9) Stems densely spreading villous; peduncle 5-14 cm   15 Hemiboea mollifolia
+ Stems glabrous to sparsely pilose toward apex; peduncle 0.2-2.1(-4) cm.   (11)
11 (10) Calyx segments 5-8 mm; style glabrous; leaf blade sparsely pilose on both surfaces, base cuneate to narrowly so; lateral veins 4-6 on each side of midrib   5 Hemiboea gracilis
+ Calyx segments 11-14 mm; style sparsely glandular puberulent or sparsely glandular; leaf blade adaxially pubescent, abaxially glabrous to sparsely pubescent, base often unequal, cuneate to rounded; lateral veins 5-11 on each side of midrib.   (12)
12 (11) Involucre sparsely pubescent outside; corolla ca. 2.1 cm; capsule ca. 1.9 cm; leaf blade margin entire; calyx segments narrowly lanceolate, ca. 2.5 mm wide   13 Hemiboea glandulosa
+ Involucre glabrous outside; corolla 3.7-4.4 cm; capsule ca. 3.5 cm; leaf blade margin conspicuously serrate to nearly entire; calyx segments lanceolate, 4-5 mm wide   14 Hemiboea fangii
13 (5) Peduncle sparsely pilose to densely pubescent.   (14)
+ Peduncle glabrous.   (17)
14 (13) Peduncle 0.2-1.2 cm; leaf blade sparsely pilose on both surfaces   5 Hemiboea gracilis
+ Peduncle 1.5-5 cm; leaf blade adaxially puberulent to pubescent, abaxially nearly glabrous to pubescent.   (15)
15 (14) Stems glabrous, much branched; leaf blade 5-8 X 2.3-4.2 cm, abaxially nearly glabrous, apex acute to abruptly acuminate   10 Hemiboea pingbianensis
+ Stems glabrescent to densely pubescent, unbranched; leaf blade 5-20 X (3-)5-10.5 cm, abaxially pubescent to sparsely pubescent, apex acute.   (16)
16 (15) Cymes 1-3-flowered; stems 5-22.5 cm with 2-5 nodes; petiole (0.5-)1-3.6 cm; leaf blade 5-12 X (3-)5-8.8 cm, without sclereids; corolla pink with purple spots, 2.1-4.1 cm; staminodes 2, 3.5-6 mm; pistil 2.1-2.5 cm   11 Hemiboea subacaulis
+ Cymes 6- or more flowered; stems ca. 40 cm with ca. 10 nodes; petiole 3-6 cm; leaf blade 14-20 X 7-10.5 cm, with vermiform sclereids surrounding vascular bundles; corolla white with purple spots, ca. 2.2 cm; staminodes 3, 0.6-3.5 mm; pistil ca. 1.4 cm   12 Hemiboea parviflora
17 (13) Calyx 2-lipped, adaxial lip 2-lobed, divided to base, abaxial lip 3-lobed, divided to middle or to apical third; stems much branched   8 Hemiboea latisepala
+ Calyx divided to base, segments equal; stems branched or unbranched.   (18)
18 (17) Calyx segments 15-20 mm.   (19)
+ Calyx segments 5-12 mm.   (20)
19 (18) Stems not spotted, nodes many; corolla glabrous outside; anthers ca. 2 mm; style glabrous   2 Hemiboea longisepala
+ Stems sparsely purple-brown spotted, nodes 3 or 4; corolla sparsely glandular puberulent outside; anthers 3.5-4 mm; style sparsely glandular pubescent   6 Hemiboea omeiensis
20 (18) Corolla glabrous outside.   (21)
+ Corolla sparsely glandular puberulent outside.   (22)
21 (20) Pistil ca. 3.2 cm; corolla pink with purple spots, tube 2.1-2.3 cm, abaxial lip 11-12 mm; petiole 0.5-2 cm; leaf blade entire, without sclereids; involucre ca. 1.3 cm in diam.   1 Hemiboea integra
+ Pistil ca. 1.5-2 cm; corolla white with purple spots, tube 3-3.8 cm, abaxial lip ca. 7 mm; petiole 2-6 cm; leaf blade margin weakly serrate to nearly entire, with vermiform sclereids surrounding vascular bundles; involucre 2-2.5 cm in diam.   3 Hemiboea bicornuta
22 (20) Leaf blade sparsely pilose; cymes 1-3-flowered; corolla pink to purple or bluish with darker or yellow lines and spots   5 Hemiboea gracilis
+ Leaf blade glabrous to pubescent; cymes (1-)3-12-flowered; corolla pale yellow to white with purple spots.   (23)
23 (22) Calyx segments 3-4.5 mm wide; pistil 3-4 cm; abaxial corolla lip 6-9 mm; leaf blade base usually slightly unequal, cuneate to attenuate and decurrent, sometimes touching at node   7 Hemiboea subcapitata
+ Calyx segments 0.2-2 mm wide; pistil 1.7-2.5 cm; abaxial corolla lip 7-15 mm; leaf blade base slightly oblique, cuneate to slightly attenuate.   (24)
24 (23) Leaf blade with vermiform sclereids surrounding vascular bundles; lateral veins 4-14 on each side of midrib; peduncle 0.5-6.5(-10) cm   4 Hemiboea cavaleriei
+ Leaf blade with astrosclereids scattered in mesophyll; lateral veins 5 or 6 on each side of midrib; peduncle 7-10 cm   9 Hemiboea longzhouensis

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