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FOC | Family List | FOC Vol. 16 | Asclepiadaceae

25. Hoya R. Brown, Prodr. 459. 1810.

球兰属 qiu lan shu

Subshrubs or lianas, epiphytic or epilithic, often twining or climbing by adventitious roots. Leaves opposite, fleshy, papery, or membranous. Inflorescences extra-axillary or sometimes terminal, umbel-like, producing a succession of globose or flat-top-ped clusters; old rachis thick, with contiguous pedicel scars. Calyx small, with basal glands. Corolla fleshy, rotate, reflexed, [or campanulate]; lobes valvate, often densely hairy or scurfy inside. Corona lobes 5, depressed patent or adnate vertically to gynostegium, fleshy, glossy when dried, inner angle a tooth incumbent on anthers, outer angle rounded or acute; margin revolute, longitudinally grooved basally. Stamens short connate, apical membrane of anthers appressed against stigma head; pollinia 2 per pollinarium, oblong, erect, with raised, translucent margin. Stigma head discoid, rounded or subapiculate. Follicles often solitary, cylindric-fusiform.

At least 100 species: SE Asia to Oceania; 32 species in China.


1 Leaves 3-veined from base   2 Hoya pottsii
+ Leaves pinnately veined, or veins obscure.   (2)
       
2 (1) Inflorescences 4- or 5-flowered, terminal; leaves 1-1.5 cm   32 Hoya chinghungensis
+ Inflorescences many flowered, usually extra-axillary, rarely terminal (H. lii); leaves more than 2 cm.   (3)
       
3 (2) Outer angle of corona lobes rounded.   (4)
+ Outer angle of corona lobes acute.   (13)
       
4 (3) Leaves villous or pubescent.   (5)
+ Leaves glabrous.   (6)
       
5 (4) Leaf blade broadly oblong or long quadrate, villous, apex mucronate; corolla lobes as long as tube   23 Hoya globulosa
+ Leaf blade elliptic, oblong, elliptic-lanceolate, or rarely suborbicular, pubescent, apex short acuminate to obtuse; corolla lobes longer than tube   24 Hoya lyi
       
6 (4) Leaf blade orbicular to widest above middle, 3-6 cm wide.   (7)
+ Leaf blade of other shapes, if more than 3 cm wide then widest below middle.   (9)
       
7 (6) Leaf blade spatulate to oblanceolate, 10-21 cm, lateral veins 10 or more pairs; flower clusters flat topped; corolla white with purple spots   21 Hoya radicalis
+ Leaf blade obovate to orbicular, 4-8 cm, lateral veins ca. 4 pairs or obscure; flower clusters globose; corolla uniformly white.   (8)
       
8 (7) Apex of leaf blade retuse or concave, lateral veins obscure   20 Hoya kerrii
+ Apex of leaf blade entire, lateral veins conspicuous, ca. 4 pairs   22 Hoya liangii
       
9 (6) Leaf blades often pandurate, sometimes oblong; peduncle 0.3 cm; corolla yellow or reddish   27 Hoya pandurata
+ Leaf blade lanceolate or oblong-lanceolate to oblong-oblanceolate; peduncle 1-8 cm; corolla white.   (10)
       
10 (9) Branchlets and petiole hairy; peduncle 1-4 cm; corolla reflexed; corona lobes erect.   (11)
+ Branchlets and petiole glabrous; peduncle 5-8 cm; corolla rotate; corona lobes spreading-stellate.   (12)
       
11 (10) Branchlets and petiole setose; lateral veins of leaf blade conspicuous adaxially; inflorescences terminal   25 Hoya lii
+ Branchlets and petiole minutely puberulent at least when young; lateral veins of leaf blade ill defined adaxially; inflorescences extra-axillary   26 Hoya mekongensis
       
12 (10) Leaf blade 1.5-2 cm wide; flower clusters flat topped; corolla 1-1.5 cm in diam., villous inside   28 Hoya mengtzeensis
+ Leaf blade 2.5-4.5 cm wide; flower clusters globose; corolla ca. 3 cm in diam., obscurely puberulent inside   29 Hoya griffithii
       
13 (3) Corolla purple; inflorescences on hanging leafless branches (leaves not known)   11 Hoya commutata
+ Corolla white, pink, yellow, or yellowish; inflorescences always on normal leafy branches.   (14)
       
14 (13) Leaf blade linear, 0.4-0.7 cm wide.   (15)
+ Leaf blade not linear, 1-6.5 cm wide.   (16)
       
15 (14) Leaf blade 2.5-5 cm, hirsute abaxially; inflorescences terminal; corolla lobes glabrous   31 Hoya linearis
+ Leaf blade 10-20 cm, glabrous; inflorescences extra-axillary; corolla lobes pubescent   30 Hoya longifolia
       
16 (14) Corolla strongly reflexed; corona distinctly stalked, lobes spurred at base   1 Hoya multiflora
+ Corolla not reflexed; corona sessile, lobes not spurred.   (17)
       
17 (16) Leaf apex caudate; flowers yellow or yellowish.   (18)
+ Leaf apex obtuse to acuminate; flowers white or reddish.   (20)
       
18 (17) Leaf blade 4.5-7 cm, base rounded, lateral veins 4 or 5 pairs, obscure; peduncle ca. 0.4 cm; pedicel 1-1.2 cm; corolla lobes glabrous   5 Hoya lasiogynostegia
+ Leaf blade 9-13 cm, base cuneate, lateral veins 10-12 pairs, conspicuous; peduncle 1.5-2.5 cm; pedicel 2-2.5 cm; corolla lobes puberulent inside.   (19)
       
19 (18) Leaf blade symmetrical at base, lateral veins at right angle to midvein, uniformly colored   3 Hoya fusca
+ Leaf blade oblique at base, lateral veins obliquely ascending, midrib with pale zone on both sides   4 Hoya lipoensis
       
20 (17) Lateral veins of leaf blade obscure or obsolete.   (21)
+ Lateral veins of leaf blade conspicuous.   (26)
       
21 (20) Leaf blade oblong or widest above middle.   (22)
+ Leaf blade widest below middle.   (23)
       
22 (21) Petiole and abaxial surface of leaf blade pubescent; corolla ca. 1.2 cm in diam.   6 Hoya thomsonii
+ Petiole and leaf blade glabrous; corolla 2-4 cm in diam.   30 Hoya longifolia
       
23 (21) Pedicel longer than peduncle.   (24)
+ Pedicel shorter than or as long as peduncle.   (25)
       
24 (23) Leaf blade lanceolate to ovate-lanceolate, apex acuminate; peduncle 1-5 cm; sepals puberulent outside   7 Hoya siamica
+ Leaf blade elliptic or elliptic-oblong, apex obtuse; peduncle ca. 0.3 cm; sepals glabrous outside   8 Hoya revolubilis
       
25 (23) Corolla 0.6-0.7 cm wide, violet dotted, base conspicuously white villous inside, lobes much longer than tube   9 Hoya lacunosa
+ Corolla 1.5-2 cm wide, not violet dotted, densely papillate inside, lobes as long as tube   10 Hoya carnosa
       
26 (20) Flower clusters flat topped.   (27)
+ Flower clusters globose.   (28)
       
27 (26) Leaf blade cordate at base; petiole and abaxial side of midvein puberulent; peduncle shorter than pedicel; calyx outside and corolla puberulent   12 Hoya cordata
+ Leaf blade rounded at base; petiole and midvein glabrous; peduncle much longer than pedicel; calyx and corolla glabrous   13 Hoya nervosa
       
28 (26) Petiole up to 0.4 cm.   (29)
+ Petiole 1-3 cm.   (30)
       
29 (28) Lateral leaf veins numerous, parallel; peduncle shorter than pedicel; corolla lobes papillate inside   14 Hoya polyneura
+ Lateral leaf veins 4 or 5 pairs, not parallel; peduncle longer than pedicel; corolla mealy inside   15 Hoya dasyantha
       
30 (28) Leaves pubescent abaxially, especially along veins.   (31)
+ Leaves glabrous.   (32)
       
31 (30) Leaf blade short acuminate at apex, cuneate at base; calyx glabrous; corolla ca. 1 cm in diam.   16 Hoya silvatica
+ Leaf blade obtuse or rounded at apex, rounded or broadly cuneate at base; calyx puberulent outside; corolla ca. 1.5 cm in diam.   17 Hoya salweenica
       
32 (30) Lateral leaf veins ca. 7 pairs; peduncle to 5 cm; calyx pubescent outside; corolla densely pale scurfy inside   18 Hoya fungii
+ Lateral leaf veins ca. 4 pairs; peduncle ca. 2 cm; calyx glabrous; corolla sparsely pubescent inside   19 Hoya ovalifolia

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