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FOC | Family List | FOC Vol. 18 | Gesneriaceae

7. Isometrum Craib, Notes Roy. Bot. Gard. Edinburgh. 11: 250. 1919.

金盏苣苔属 jin zhun ju tai shu

Herbs, perennial, epipetric or terrestrial, rhizomatous, stemless. Leaves many, basal; leaf blade puberulent to villous, tomentose, or glabrescent, base narrowly cuneate to cordate. Inflorescences cymes, rarely umbel-like, lax or rarely dense, axillary, 2- to many flowered; bracts 2(or 3), opposite, rarely whorled. Calyx actinomorphic, 5-sect from base; segments equal. Corolla white to purple or pink, rarely yellow, zygomorphic, rarely actinomorphic, inside glabrous or sparsely puberulent, rarely pilose on lobes; tube campanulate, cylindric, rarely urceolate or salverform, not swollen, 1-4 X longer than limb, 3-11 mm in diam.; limb slightly 2-lipped; adaxial lip 2-lobed from near middle, shorter than to nearly equalling abaxial lip; abaxial lip 3-sect from base, lobes equal or subequal, apex rounded, sometimes crenulate. Stamens 4, adnate to corolla tube near base, rarely to or above middle, included; anthers basifixed, coherent in pairs, rarely free, thecae divergent at base, not or rarely confluent at apex, dehiscing longitudinally from arcuate slits; connective not projecting; staminode 1, adnate to adaxial side of corolla tube. Disc ringlike. Ovary narrowly oblong to oblong or ovoid, 1-loculed; placentas 2, parietal, projecting inward, 2-cleft. Stigmas 2, equal, oblate, undivided. Capsule straight in relation to pedicel, narrowly oblong to oblanceolate, much longer than calyx, dehiscing loculicidally to base; valves 2, straight, not twisted. Seeds unappendaged.

* Fourteen species: endemic to China.

1 Corolla tube 1-2 X longer than broad; pistil glabrous.   (2)
+ Corolla tube 1.5-5 X longer than broad; pistil glabrous or puberulent to pubescent.   (5)
2 (1) Corolla puberulent outside; leaf blade adaxially densely brown villous, abaxially densely brown long woolly   2 Isometrum glandulosum
+ Corolla glabrous outside; leaf blade adaxially puberulent, puberulent-strigillose, or sparsely brown villous, abaxially puberulent to pubescent or sparsely villous.   (3)
3 (2) Stamens free, anther thecae confluent; leaf blade margin lobed, lobes triangular, deeply serrate to dentate; lateral veins 7-13 on each side of midrib; bract and sepal margin sparsely dentate   1 Isometrum eximium
+ Stamens coherent in pairs, anther thecae not confluent; leaf blade margin serrate, at least near apex; lateral veins 3-5 on each side of midrib; bract and sepal margin entire.   (4)
4 (3) Petiole to 1.5 cm; leaf blade base cuneate; peduncle villous, glabrescent; corolla tube campanulate; pistil 6-7 mm   3 Isometrum fargesii
+ Petiole to 8.3 cm; leaf blade base cordate; peduncle puberulent and glandular puberulent; corolla tube urceolate; pistil 3.8-4 mm   4 Isometrum nanchuanicum
5 (1) Leaf blade margin lobulate.   (6)
+ Leaf blade margin subentire or sinuous to coarsely crenate-dentate.   (10)
6 (5) Corolla tube campanulate-tubular to narrowly campanulate; petiole to 2 cm.   (7)
+ Corolla tube nearly cylindric; petiole to 4 cm.   (8)
7 (6) Leaf blade 1-1.8 cm wide; corolla 1-1.5 cm, adaxial lip lobes ca. 2 X 5 mm; stamens 6-8.5 mm, filaments sparsely puberulent to glabrous; pistil glabrous   6 Isometrum farreri
+ Leaf blade 2-3 cm wide; corolla 1.6-2 cm, adaxial lip lobes 4-6 X 3-4 mm; stamens 3-4 mm, filaments glabrous; pistil pubescent   7 Isometrum giraldii
8 (6) Leaf blade oblanceolate, 1.5-5 X 0.8-1.5 cm, margin lobes subentire to shallowly crenulate; lateral veins 3-5 on each side of midrib; peduncle 4-6 cm; bract 1.8-2 mm   14 Isometrum pinnatilobatum
+ Leaf blade narrowly elliptic to ovate or elliptic, 2.7-10.5 X 2.3-5.5 cm, margin lobes crenate- dentate or crenate; lateral veins 5-8 on each side of midrib; peduncle 4-20 cm; bract 3-5 mm.   (9)
9 (8) Corolla tube 2-2.7 mm in diam., adaxial lip ca. 5 mm, abaxial lip ca. 5.5 mm; stamens 3.5-4 mm; pistil glabrous; leaf blade base cuneate   12 Isometrum primuliflorum
+ Corolla tube 3-3.5 mm in diam., adaxial lip ca. 3.5 mm, abaxial lip ca. 3 mm; stamens 5-6 mm; pistil glandular puberulent; leaf blade base narrowly cuneate   13 Isometrum sichuanicum
10 (5) Petiole to 8 cm.   (11)
+ Leaves sessile or petiole to 2 cm.   (13)
11 (10) Corolla white, minutely pilose outside; leaf blade 3-4.5 cm   5 Isometrum leucanthum
+ Corolla pink to purple-red or pale purple to purple, glandular puberulent or sparsely glandular pubescent, sometimes only basally; leaf blade (2-)4-14 cm.   (12)
12 (11) Petiole densely brown villous to lanate; leaf blade abaxially rust-brown villous, base narrowly cuneate to truncate; lateral veins 6-10 on each side of midrib; pedicel 1-2.2 cm; corolla 0.8-2 cm, tube narrowly urceolate, ca. 3 mm in diam., lobes 1-2 mm; stamens 4-5 mm; pistil glabrous   9 Isometrum lancifolium
+ Petiole densely white pubescent; leaf blade abaxially pubescent, base rounded to cordate; lateral veins 4-6 on each side of midrib; pedicel 4-7 mm; corolla 2.3-3.7 cm, tube campanulate-tubular, 7-11 mm in diam.; lobes 4-7(-11) mm; stamens 7-10 mm; pistil sparsely pubescent near apex   11 Isometrum lungshengense
13 (10) Calyx segments 4-6 mm; leaf blade base narrowly cuneate to subtruncate   7 Isometrum giraldii
+ Calyx segments 2-3.5 mm; leaf blade base narrowly cuneate or cuneate.   (14)
14 (13) Leaf blade sessile, 1.5-3.7 X 1-1.8 cm, margin irregularly dentate above middle; pistil ca. 4 mm   10 Isometrum villosum
+ Petiole to 2 cm; leaf blade 3-8 X 1.5-3.5 cm, margin crenate to lobulate; pistil 9-10 mm.   (15)
15 (14) Petiole to 1 cm; leaf blade both surfaces gray pubescent, veins abaxially brown villous; lateral veins abaxially prominent; peduncle 4-17 cm; bracts 3.5-5 mm; adaxial corolla lobes ca. 2 mm, abaxial lobes ca. 4 mm; stamens 6-8.5 mm; pistil glabrous   6 Isometrum farreri
+ Petiole to 2 cm; leaf blade adaxially gray appressed pubescent, abaxially brown villous; lateral veins indistinct; peduncle 5.5-10 cm; bracts ca. 2.5 mm; all corolla lobes 3-5 mm; stamens 3.5-4.5 mm; pistil puberulent to subglabrous   8 Isometrum crenatum

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