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FOC | Family List | FOC Vol. 19 | Acanthaceae

30. Lepidagathis Willdenow, Sp. Pl. 3: 400. 1800.

鳞花草属 lin hua cao shu

Authors: Jia-qi Hu & Thomas F. Daniel

Herbs [or shrubs], perennial, with cystoliths, often anisophyllous. Leaves opposite; leaf blade margin entire or crenate-serrate. Inflorescences dense spikes or cymes condensed into globose heads, sometimes clustered; bracts and bracteoles present. Calyx 5-lobed; lobes heteromorphic; posterior lobe usually largest; lateral 2 lobes smallest; anterior 2 lobes usually connate to a greater degree than other lobes. Corolla small; tube usually expanded into a distinct throat near middle; limb 2-lipped; lower lip 3-lobed, patent, usually spotted, lobes subequal or middle one slightly larger; upper lip erect or slightly patent, entire to 2-lobed; lobes quincuncial in bud. Stamens 4, didynamous [or with one pair reduced or absent], usually included in corolla tube; filaments short; anthers 2-thecous [or with one pair of anthers sometimes 1-thecous]; thecae oblong, parallel or one lower and somewhat divergent, base muticous or spurred. Ovary with 2 ovules per locule; stigma subcapitate, entire or slightly 2-cleft. Capsule subellipsoid to oblong to linear-conical, lacking a stalk, to 4-seeded; retinacula present. Seeds discoid, pubescent with hygroscopic trichomes.

About 100 species: tropical to subtropical regions worldwide (mostly paleotropical); seven species (three endemic) in China.

1 Inflorescences globose or ovoid heads to 1 cm; leaves sessile to subsessile; leaf blade usually less than 2 cm, base truncate to subcordate.   6 L. secunda
+ Inflorescences ± elongate and secund spikes to 5.5 cm; leaves usually petiolate; leaf blade to 14.5 cm, base cuneate to attenuate and usually decurrent onto petiole (in L. inaequalis abruptly truncate to rounded and decurrent onto petiole)   (2)
2 (1) Leaf blade linear to linear-lanceolate, (4.6-)6.5-16 × as long as wide.   7 L. stenophylla
+ Leaf blade ovate to elliptic to subcircular to obovate, 1.6-4.5(-5.6) × as long as wide (in L. fasciculata, sometimes only distalmost 1 or 2 nodes with leaf blade linear to linear-lanceolate and 6-9.5 × as long as wide and others ovate to elliptic and 1.7-3 × as long as wide)   (3)
3 (2) Anterior calyx lobes connate at base for 10%-25% of their length; bracts (1 or)3(or 5)-veined   (4)
+ Anterior calyx lobes connate at base for 30%-60% of their length; bracts 1(or 3)-veined   (5)
4 (3) Corolla pinkish; bracts broadly ovate to ovate-oblong, 2.5-4.4 mm wide; posterior calyx lobe 3-veined; distinct segments of anterior calyx lobe oblanceolate to linear, 6-6.5 mm; capsule 4.5-5 mm, pubescent distally.   1 L. fasciculata
+ Corolla white; bracts narrowly lanceolate, 1-1.5 mm wide; posterior calyx lobe 5-veined; distinct segments of anterior calyx lobe narrowly lanceolate, 9-10 mm; capsule 6-6.5 mm, pubescent throughout.   3 L. hainanensis
5 (3) Leaf blade 1-4(-6) cm, 1.6-2 × as long as wide, base abruptly rounded to truncate and decurrent onto petiole; plants decumbent, not conspicuously woody at base.   4 L. inaequalis
+ Leaf blade to 14.5 cm, usually more than 2 × as long as wide, base cuneate to long attenuate and decurrent onto petiole; plants not decumbent, conspicuously woody at base   (6)
6 (5) Calyx 5-6 mm.   2 L. formosensis
+ Calyx 8-10 mm.   5 L. incurva

Lower Taxa


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