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FOC | Family List | FOC Vol. 22 | Poaceae

98. Leymus Hochstetter, Flora. 31: 118. 1848.

赖草属 lai cao shu

Aneurolepidium Nevski.

Herbs perennial. Plants usually with rhizomes, rarely without. Culms usually erect. Leaf sheath split almost to base; ligule leathery-membranous; auricles lanceolate or crescent-shaped; leaf blade usually glaucous, usually rolled, rarely flat, stiff, harsh. Inflorescence spikelike, linear, rarely oblong-ovate; rachis tough. Spikelets in regular rows of (1 or)2 or 3(–6) per node, sessile, all similar, with (1–)3–7 florets; rachilla disarticulating above glumes and between florets. Glumes opposite or side-by-side, linear to lanceolate, 1–3(–5)-veined, veins not raised (except in L. mollis), not keeled or keeled almost to base, apex acute to shortly awned. Lemma 3–7-veined, abaxially not keeled or keeled only at apex, apex acute to shortly awned; callus obtuse, triangular or rounded. Lodicules lanceolate to ovate, usually entire. Caryopsis adherent to lemma.

About 50 species:temperate regions of the N hemisphere; 24 species (11 endemic) in China.

The authors have seen no specimens of Leymus pubinodis (Keng) A. Löve (Repert. Spec. Nov. Regni Veg. 95: 481. 1984, "pubinodes"; Elymus pubinodis Keng, Sunyatsenia 6: 85. 1941), described from Xizang.


1 Spikelets (3 or)4–6 per node   (2)
+ Spikelets 1–3(or 4) per node   (5)
       
2 (1) Spikelets with 3–5 florets; first lemma 15–20 mm.   1 L. racemosus
+ Spikelets with 5–10 florets; first lemma 8–13 mm   (3)
       
3 (2) Leaf blade 5–15 × 0.5–0.8 cm; spike oblong-ovate, 5–9 cm.   4 L. ovatus
+ Leaf blade 15–42 cm; spike linear, 12–25 cm   (4)
       
4 (3) Glumes 3-veined, unequal, proximal one 10–12 mm, distal one 13–16 mm; spikelets 3–5 per node.   2 L. pseudoracemosus
+ Glumes 1-veined, subequal, 10–13 mm; spikelets 4–6(–11) per node.   3 L. crassiusculus
       
5 (1) Spikelets usually 1 per node, rarely 2 or 3 per node below middle of spike   (6)
+ Spikelets 1–3(or 4) per node   (12)
       
6 (5) Glumes lanceolate or linear-lanceolate, 3-veined   (7)
+ Glumes subulate-linear or narrowly lanceolate, veinless or 1-veined   (9)
       
7 (6) Palea much shorter than lemma, apex emarginate; spikelets with 2 or 3 florets.   18 L. pishanicus
+ Palea slightly shorter than or equaling lemma; spikelets with 4–8 florets   (8)
       
8 (7) Glumes 10–15 mm, margin membranous and ciliate.   19 L. altus
+ Glumes 6–7.5 mm, margin membranous but not not ciliate.   20 L. obvipodus
       
9 (6) Lemma puberulent, or if smooth and glabrous then margin ciliate; leaf blade 0.1–0.3 cm wide.   21 L. ruoqiangensis
+ Lemma smooth, glabrous; leaf blade 0.2–2.5 cm wide   (10)
       
10 (9) Leaf blade to 2.5 cm wide; glumes subulate-linear.   23 L. aemulans
+ Leaf blade 0.2–0.6 cm wide, glumes linear-lanceolate   (11)
       
11 (10) Spikelets with 3 or 4 florets.   22 L. arjinshanicus
+ Spikelets with 4–7 florets.   24 L. ramosus
       
12 (5) Glumes equaling or slightly longer than spikelet   (13)
+ Glumes shorter than spikelet   (16)
       
13 (12) Glumes wholly covering lemma or covering only base of first lemma   (14)
+ Glumes not covering lemma   (15)
       
14 (13) Spikelets 2 per node; glumes wholly covering lemma.   10 L. karelinii
+ Spikelets (1 or)2 or 3 per node; glumes covering only base of first lemma.   11 L. mollis
       
15 (13) Ligule ca. 0.5 mm; rachis internodes 3–6 mm.   12 L. paboanus
+ Ligule 2–3 mm; rachis internodes 6–20 mm.   13 L. tianschanicus
       
16 (12) Glumes oblong-lanceolate, apex with awn 4–5 mm.   5 L. aristiglumis
+ Glumes not as above   (17)
       
17 (16) Glumes covering base of first lemma   (18)
+ Glumes not covering base of first lemma   (21)
       
18 (17) First lemma 7–9 mm including apical point   (19)
+ First lemma 10–14 mm   (20)
       
19 (18) Culms 14–35 cm tall; glumes narrowly lanceolate, 5–9 mm, margin not ciliate.   7 L. yiunensis
+ Culms 60–150 cm tall; glumes linear-lanceolate, 9–11 mm, distal margin sparsely ciliate.   8 L. pendulus
       
20 (18) Spikelets with 2 or 3 florets.   6 L. angustus
+ Spikelets with 5–9 florets.   9 L. shanxiensis
       
21 (17) Glumes 3-veined; palea equaling lemma   (22)
+ Glumes 1-veined or nearly veinless; palea shorter than or subequaling lemma   (23)
       
22 (21) Callus of lemma villous; glumes linear-lanceolate.   15 L. secalinus
+ Callus of lemma smooth; glumes subulate.   16 L. chinensis
       
23 (21) Callus of lemma puberulent; palea shorter than lemma; first lemma 5–8 mm.   14 L. multicaulis
+ Callus of lemma villous; palea subequaling lemma; first lemma 9–10 mm.   17 L. flexus

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