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59. Libanotis Haller ex Zinn, Cat. Pl. Hort. Gott. 226. 1757, nom. cons., not Hill (1756).
岩风属 yan feng shu
She Menglan (佘孟兰 Sheh Meng-lan); Michael G. Pimenov, Eugene V. Kljuykov, Mark F. Watson
Herbs rarely subshrubs, perennial, stout, sometimes small, rarely acaulescent. Taproot conic, unbranched, caudex simple, rarely branched, woody. Stem often strongly angled and fluted, base densely clothed with fibrous leaf remains. Basal leaves 1–4-pinnate or 1–4-pinnatisect; ultimate segments linear, ovate or lanceolate, entire or lobed. Umbels compound, terminal and lateral; bracts few to numerous or absent; rays numerous to few; bracteoles several, linear or lanceolate. Calyx teeth conspicuous, linear, triangular or elliptic. Petals white, rarely pinkish, ovate or obcordate, apex narrow, inflexed. Stylopodium low-conic, margins often undulate at the base. Fruit ovoid or oblong, slightly to moderately dorsally compressed; dorsal ribs filiform, low or prominent, acute-ridged, lateral ribs sometimes slightly broader; vittae 1–2(–3) in each furrow, 2–4 (rarely 6–8) on commissure. Seed face plane. Carpophore entire or 2-parted.
About 30 species: Asia, Europe; 18 species (eight endemic) in China.
The taxonomic position of Libanotis and related genera continues to be controversial. Since its establishment authors have either accepted it as an independent genus or regarded it as a subgenus or a section of Seseli. The conspicuous calyx teeth, well-developed bracts and bracteoles, and almost always pubescent fruit seem to distinguish it from Seseli, but more material is needed for a taxonomic study to resolve this complex situation. Among the present co-authors, Pimenov and Kljuykov favor a classification where Eriocycla and Libanotis are included within an expanded, polymorphic Seseli, whereas She and Watson prefer to retain these as separate genera pending further research. As is usual in Flora of China, where there are differences of taxonomic opinion, the Chinese view is adopted for the Flora and a summary of the alternative classification is provided, in this case at the end of the Seseli account.
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| 1 |
Plants acaulescent, 2–10 cm; leaves in rosette; fruit densely papillose-scaly. |
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(2) |
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Plants caulescent, 10–130 cm; leaves not in rosette; fruit glabrous or variously hairy, not papillose-scaly. |
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(3) |
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| 2 (1) |
Leaf sheaths broadly ovate; fruit vittae 2–3 in each furrow, 4–6 on commissure. |
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17 Libanotis acaulis |
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Leaf sheaths narrowly lanceolate; fruit vittae solitary in each furrow, 2 on commissure. |
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18 Libanotis depressa |
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| 3 (1) |
Petals abaxially glabrous or sparsely pubescent; caudex short; stem strongly angled and fluted; leaves papery, matt. |
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(4) |
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Petals abaxially pubescent or villous; caudex stout, elongate and exposed; stem terete; leaves thick or
coriaceous, shiny. |
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(8) |
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| 4 (3) |
Stem hollow; bracteoles longer than umbellules. |
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12 Libanotis condensata |
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Stem rigid, solid; bracteoles shorter than umbellules. |
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(5) |
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| 5 (4) |
Fruit vittae 3–4 in each furrow, 6–8 on commissure. |
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13 Libanotis seseloides |
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Fruit vittae solitary in each furrow, 2–4 on commissure. |
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(6) |
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| 6 (5) |
Leaf blade narrowly elliptic, 3-pinnatisect, ultimate segments linear, 1–2 × 0.4–0.6 mm. |
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14 Libanotis incana |
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Leaf blade oblong, 2-pinnatisect, ultimate segments ovate rhombic or linear, 8–15 × 1.2–7 mm. |
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(7) |
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| 7 (6) |
Stem solitary; rays 35–50. |
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15 Libanotis sibirica |
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Stems usually several; rays 15–25. |
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16 Libanotis schrenkiana |
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| 8 (3) |
Rays 30–50; fruit ribs acute, prominent. |
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1 Libanotis buchtormensis |
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Rays 2–25; fruit ribs obtuse or rounded. |
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(9) |
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| 9 (8) |
Stem suberect, branches very slender, diffusely spreading; rays 2–4. |
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4 Libanotis lanzhouensis |
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Stem erect, branches stout and ascending-spreading; rays 4–25 (except L. laticalycina). |
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(10) |
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| 10 (9) |
Leaf pinnae long-petiolulate. |
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(11) |
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Leaf pinnae sessile or near so. |
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(13) |
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| 11 (10) |
Ultimate leaf segments narrow-lanceolate or elliptic-lanceolate, entire. |
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6 Libanotis lancifolia |
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Ultimate leaf segments obovate, usually dentate or 2–3-lobed. |
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(12) |
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| 12 (11) |
Rays 2–4, short, 1.5–4 mm. |
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7 Libanotis laticalycina |
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Rays 5–9, long, 15–30 mm. |
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8 Libanotis jinanensis |
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| 13 (10) |
Ultimate leaf segments linear, entire. |
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2 Libanotis iliensis |
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Ultimate leaf segments linear-lanceolate or ovate, toothed or shallowly lobed. |
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(14) |
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| 14 (13) |
Plants 40–90 cm, usually subshrubs. |
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(15) |
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Plants 10–40(–60) cm, entirely herbaceous. |
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(16) |
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| 15 (14) |
Plants grayish pubescent throughout. |
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5 Libanotis spodotrichoma |
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Plants white pubescent throughout. |
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3 Libanotis wannienchun |
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| 16 (14) |
Rays 4–8; fruit densely lanate with gray hairs. |
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9 Libanotis eriocarpa |
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Rays 6–15; fruit densely hispid or shortly pubescent, not lanate. |
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(17) |
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| 17 (16) |
Ultimate leaf segments linear-lanceolate, 1–5 × 0.5–1.5 mm; marginal fruit ribs slightly larger than dorsal; vittae solitary in each furrow, 2 on commissure. |
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10 Libanotis abolinii |
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Ultimate leaf segments oblong to linear-lanceolate, 5–40 × 3–7 mm; marginal fruit ribs considerably larger than dorsal; vittae (1–)2–3 in each furrow,
(2–)4–6 on commissure. |
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11 Libanotis grubovii |
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List of lower taxa
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