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FOC | Family List | FOC Vol. 13 | Melastomataceae

19. Medinilla Gaudichaud-Beaupré ex Candolle, Prodr. 3: 167. 1828.

酸脚杆属 suan jiao gan shu

Pseudodissochaeta M. P. Nayar.

Small trees, or erect or scandent shrubs, terrestrial, epiphytic, or rarely climbing. Stems 4-sided or terete, sometimes succulent or corky. Leaves opposite or verticillate, petiolate or sessile; leaf blade usually glabrous, margin entire or dentate. Inflorescences terminal or axillary inserted on leafless stems or at nodes of root stock, cymose or cymose paniculate; bracts small, caducous. Flowers 4(-6)-merous, pedicellate and sometimes bracteolate. Hypanthium cup-shaped, funnel-shaped, campanulate, or tubular. Calyx lobes conspicuous or inconspicuous, apex apiculate. Petals obovate, ovate, or suborbicular, sometimes oblique. Stamens 2 × as many as petals, whorls equal or slightly unequal in shape and length; anthers linear, lanceolate, or oblong, base tuberculate or appendaged, apex beaked, dehiscent by a single pore; connective slightly decurrent forming a spur. Ovary inferior, ovate, apex truncate or with a membranous crown, sometimes septate. Berry globular, ovate, or often urceolate, apex indehiscent. Seeds numerous, obovate to shortly cuneate, small, glabrous or tuberculate.

Between 300 and 400 species: tropical Africa, Asia, and Pacific islands; 11 species (five endemic) in China.

In China Pseudodissochaeta has been treated as a synonym of Medinilla (the species concerned are M. assamica, M. lanceata, and M. septentrionalis), whereas Pseudodissochaeta has also been treated as a separate genus (Fl. Thailand 7(3): 475. 2001). Unpublished molecular data indicate that at least M. septentrionalis (P. septentrionalis) is not close to Medinilla. The fact that the Melastomataceae specialists W. W. Smith, H. L. Li, Kraenzlin, Guillaumin, Nayar, and Chen Cheih allocated the three Chinese species of Pseudodissochaeta to six different genera (Allomorphia, Anplectrum A. Gray, Diplectria, Medinilla, Oritrephes Ridley, Pseudodissochaeta) indicates the difficulty of determining their true affinities from morphological data. Additional molecular data are needed to resolve the problem.


1 Leaves 4-8-verticillate, leaf blade oblong, 9-12 × 2.5-3 cm.   7 M. hayatana
+ Leaves opposite, leaf blade lanceolate, elliptic, or obovate, but not oblong   (2)
       
2 (1) Branchlets conspicuously long setose at leaf nodes, seta to 8 mm.   6 M. formosana
+ Branchlets not setose at leaf nodes   (3)
       
3 (2) Leaf blade elliptic, (2-)6-8.5 × 1-2.8 cm   (4)
+ Leaf blade larger than above   (5)
       
4 (3) Leaves and branchlets not succulent, leaves drying thin-papery; branchlets terete, with thin not corky bark.   1 M. septentrionalis
+ Leaves and branchlets succulent, drying subleathery; branchlets angular, with thickly corky bark.   4 M. nana
       
5 (3) Inflorescences 15-30 cm, many-flowered; leaf blade abaxially puberulous, furfuraceous, or finely strigose   (6)
+ Inflorescences shorter than above, usually few-flowered; leaf blade abaxially glabrous   (7)
       
6 (5) Leaves sessile or subsessile; leaf blade 3.8-11 cm wide.   2 M. assamica
+ Leaves with a 0.8-1 cm petiole; leaf blade 3-3.5 cm wide.   3 M. lanceata
       
7 (5) Bark of branchlets dark brown with conspicuous white lenticels.   5 M. fengii
+ Bark of branchlets usually pale tan (not dark brown), rarely conspicuously lenticellate   (8)
       
8 (7) Leaves 3-5-verticillate, leaf blade obovate, 2-3.5(-5) × 1.1-1.5(-2.8) cm   8 M. arboricola
+ Leaves opposite, leaf blade elliptic, 6-23 × 2-11 cm   (9)
       
9 (8) Inflorescences terminal or lateral cymes, peduncle 2-3 cm.   9 M. himalayana
+ Inflorescences lateral cymes only, usually on leafless branchlets, peduncle 0.5-1 cm   (10)
       
10 (9) Leaf blade base subrounded, not decurrent.   10 M. rubicunda
+ Leaf blade base cuneate and slightly decurrent.   11 M. petelotii

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