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115. Nannoglottis Maximowicz, Bull. Acad. Imp. Sci. Saint-Pétersbourg. 27: 480. 1882.

毛冠菊属 mao guan ju shu

Authors: Yilin Chen & Luc Brouillet

Stereosanthus Franchet; Vierhapperia Handel-Mazzetti.

Herbs, perennial, rarely subshrubs; rhizome woody; herbage sometimes or partly long white lanate or cottony, stipitate glandular. Leaves alternate, basal and lower cauline long or shortly petiolate, blade lanceolate, ovate-lanceolate, oblong, orbicular, or cordate, margin coarsely serrate, mid and upper cauline sessile, often decurrent. Capitula ± large, few to numerous in corymbiform synflorescences, rarely solitary, radiate. Involucre hemispheric or cup-shaped; phyllaries 2-4-seriate, unequal to equal, herbaceous, rarely papery, multiveined, margins of inner narrowly scarious, outer glandular or long hairy. Receptacles flat or slightly convex, ± alveolate, naked. Ray florets outer with lamina yellow, reddish, or brownish, surpassing or ± equaling phyllaries, inner 2- or 3-seriate, rarely few or absent, tubular, elaminate, shorter than styles; disk functionally male, yellow, narrowly funnelform, 5-lobed, lobes lanceolate to linear-lanceolate, often glandular; anther bases obtuse or acute; style branches of ray florets linear-lanceolate, long, of disk ± appressed together, lanceolate, ± short, densely hairy (outer), without receptive papillae. Achenes brown, oblong, sparsely strigose, pale 8-10-veined. Pappus of few short, thin bristles (sometimes absent), and many long, coarse or thin, long-barbellate bristles, those of disk sometimes few, short, thin.

Nine species: Bhutan, China, India, Nepal; nine species (six endemic) in China.

J. Q. Liu et al. (Molec. Phylogen. Evol. 23: 307-325. 2002) produced a molecular phylogeny of Nannoglottis, showing it is one of the earliest diverging lineages of Astereae. Nannoglottis ravida is the earliest diverging member of the genus and has been segregated under N. sect. Monocephala T. G. Gao & Y. L. Chen from all other species of the genus.

1 Subshrubs; capitula solitary; ray florets yellow; leaves abaxially densely cottony.   1 N. ravida
+ Perennial herbs; capitula numerous in corymbiform synflorescences; ray florets yellow, brownish, or reddish; leaves abaxially ± densely cottony or arachnoid, or glandular   (2)
2 (1) Ray florets brownish or reddish, lamina slightly exceeding involucre   (3)
+ Ray florets yellow, lamina distinctly exceeding involucre   (5)
3 (2) Basal leaves cordate, long winged petiolate.   9 N. yunnanensis
+ Basal leaves ovate-lanceolate, attenuate at base   (4)
4 (3) Mid cauline leaves base abruptly narrowed into short, broadly winged petiole, margin denticulate, abaxially stipitate glandular.   8 N. hieraciophylla
+ Mid cauline leaves base attenuate, margin dentate, abaxially stipitate glandular, sparsely arachnoid.   7 N. carpesioides
5 (2) Basal leaves abaxially densely lanate (glands not visible), sometimes arachnoid   (6)
+ Basal leaves abaxially glandular or also arachnoid   (7)
6 (5) Basal leaves long petiolate, cordate to orbicular or broadly ovate, margin shallowly dentate, teeth obtuse or acute, mid cauline abruptly smaller; phyllaries lanceolate.   2 N. delavayi
+ Basal leaves shortly petiolate, ovate to lanceolate, margin irregularly denticulate to dentate, mid cauline developed; phyllaries ovate to lanceolate-ovate.   5 N. latisquama
7 (5) Phyllaries papery, 3- or 4-seriate, unequal, outer shorter than inner.   3 N. gynura
+ Phyllaries herbaceous, 2- or 3-seriate, equal or subequal   (8)
8 (7) Basal leaves attenuate at base, decurrent on stem, usually sessile.   4 N. hookeri
+ Basal leaves abruptly narrowed, long petiolate.   6 N. macrocarpa

Lower Taxa


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