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52. PADUS Miller, Gard. Dict. Abr, ed. 4. [999]. 1754.

稠李属 chou li shu

Gu Cuizhi (Ku Tsue-chih); Bruce Bartholomew

Trees or shrubs, deciduous, many branched. Branches unarmed. Axillary winter buds ovoid; terminal bud present. Stipules membranous, soon caducous. Leaves simple, alternate, conduplicate when young; petiole usually with 2 nectaries at apex or at base of leaf blade margin; leaf blade margin serrate, rarely entire. Inflorescences terminal on current year’s branchlet, racemose, many-flowered, base with a soon caducous involucre formed by floral bud scales; peduncle usually with leaves. Hypanthium campanulate to cup-shaped. Sepals 5. Petals 5, white. Stamens 10 or more, inserted on rim of hypanthium. Ovary superior, 1-loculed; ovules 2, collateral, pendulous. Style terminal, elongated; stigma flat. Fruit a drupe, glabrous, not glaucous, without a longitudinal groove; mesocarp succulent, not splitting when ripe; endocarp bony.

About 20 species: mostly in N temperate regions; 15 species (nine endemic) in China.

1 Racemes basally leafless; sepals persistent in fruit.   (2)
+ Racemes basally leafy (usually leafless in P. maackii and P. laxiflora); sepals soon caducous in fruit.   (4)
2 (1) Branchlets and abaxial surface of leaf blades glabrous; peduncle subglabrous to sparsely pubescent.   1 P. buergeriana
+ Branchlets pubescent and leaf blades pubescent at least on veins; peduncle densely pubescent.   (3)
3 (2) Leaf blade margin acutely serrate; inflorescence base without squamose involucral bracts.   2 P. stellipila
+ Leaf blade margin serrulate; inflorescence base with persistent squamose involucral bracts.   3 P. perulata
4 (1) Inflorescence base leafless or sometimes with 1 or 2 small bracts near base; leaves abaxially glandular.   (5)
+ Inflorescences base leafy; leaf blade abaxially non-glandular.   (6)
5 (4) Leaf blade abaxially glandular, midvein pubescent.   4 P. maackii
+ Leaf blade not glandular, midvein sericeous-villous.   5 P. laxiflora
6 (4) Peduncles and pedicels thickened, conspicuously lenticellate in fruit.   (7)
+ Peduncles and pedicels not thickened and not lenticellate in fruit.   (8)
7 (6) Branchlets and abaxial surface of leaf blades glabrous or very rarely sparsely pubescent when young.   14 P. napaulensis
+ Branchlets and abaxial surface of leaf blades pubescent.   15 P. wilsonii
8 (6) Style exserted.   6 P. grayana
+ Style not exserted (not seen in P. brunnescens or P. integrifolia).   (9)
9 (8) Style 1/2 as long as stamens; pedicel 1–1.5(–2.4) cm.   7 P. avium
+ Style nearly as long as stamens (not seen in P. brunnescens or P. integrifolia); pedicel shorter than 1 cm.   (10)
10 (9) Leaf blade margin serrulate.   13 P. cornuta
+ Leaf blade margin entire, crenulate, or serrate.   (11)
11 (10) Leaf blade margin aristate serrate; inflorescences 16–30 cm.   (12)
+ Leaf blade margin not aristate serrate; inflorescences 8–15 cm.   (13)
12 (11) Leaf blade elliptic to ovate-oblong, abaxially brown pubescent; peduncles and pedicels brown pubescent.   8 P. brunnescens
+ Leaf blade oblong to rarely elliptic or lanceolate, glabrous; peduncles and pedicels not brown pubescent.   9 P. brachypoda
13 (11) Leaf blade margin entire or very rarely apically sparsely serrate.   12 P. integrifolia
+ Leaf blade margin crenulate or appressed serrate.   (14)
14 (13) Leaf blade abaxially glabrous; branchlets glabrous or pubescent; peduncles and pedicels pubescent.   10 P. obtusata
+ Leaf blade abaxially velutinous; branchlets velutinous; peduncles and pedicels velutinous.   11 P. velutina

  • List of lower taxa


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