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41. Peristylus Blume, Bijdr. 404. 1825.
阔蕊兰属 kuo rui lan shu
Authors: Xinqi Chen, Stephan W. Gale & Phillip J. Cribb
Glossaspis Sprengel; Glossula Lindley.
Herbs, terrestrial, slender to robust, medium-sized. Tubers subglobose to ellipsoid-oblong, undivided, fleshy, neck with several slender roots. Stem erect, with 2 or 3 tubular sheaths at base, 1- to many leaved, glabrous. Leaves basal, clustered midway along stem or widely spaced, elliptic, ovate, oblanceolate, or linear, base contracted into amplexicaul sheath. Inflorescence erect, terminal, racemose, glabrous; peduncle often with 1 to several sterile bracts; rachis laxly to densely many flowered. Flowers spirally arranged, resupinate, small, sometimes spidery, green, yellowish green, or white; ovary and pedicel usually erect and held close to rachis, twisted, cylindric-fusiform, slender, usually glabrous. Sepals free; dorsal sepal erect; lateral sepals usually spreading, rarely reflexed. Petals often connivent with dorsal sepal and forming a hood, slightly fleshy; lip connate with column at base, distally 3-lobed or rarely entire, larger than sepals and petals, slightly fleshy, spurred at base, ecallose or with a fleshy callus above mouth of spur; spur short and scrotiform, or sometimes elongate and cylindric, usually much shorter than ovary. Column erect, very short, stout; anther with 2 parallel or slightly divergent locules; pollinia 2, granular-farinaceous, sectile, each attached to a viscidium via a short caudicle; viscidia often small, naked, elliptic, ovate, or suborbicular; rostellum small, with short, inconspicuous arms; stigma lobes 2, basally connate but diverging widely, convex, globose or subclavate, often adnate to base of lip; auricles 2, prominent, placed laterally at base of anther.
About 70 species: E, S, and SE Asia to New Guinea, NE Australia, and the SW Pacific islands; 19 species (five endemic) in China.
After publication of the treatment for the Flora of China, one species was newly recorded from China (see JIN Xiao-Hua et al. 2010. Acta Bot. Yunnan. 32: 331-333, i.e., Peristylus superanthus J. J. Wood).
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| 1 |
Lip entire |
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(2) |
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Lip 3-lobed |
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(4) |
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| 2 (1) |
Leaves linear, 0.2-0.3 cm wide; spur 5.5-6(-8) mm, ca. as long as ovary. |
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17 P. forrestii |
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Leaves oblong, oblong-elliptic, elliptic-lanceolate, or lanceolate, 0.7-2 cm wide; spur 0.7-1.2 mm, much shorter than ovary |
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(3) |
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| 3 (2) |
Plants 2-5-leaved; floral bracts ca. 8 mm, exceeding flower. |
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18 P. forceps |
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Plants 1-leaved; floral bracts 4-5 mm, almost as long as to slightly exceeding ovary. |
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19 P. nematocaulon |
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| 4 (1) |
Lateral lobes of lip filiform or linear, ca. 2 × as long as mid-lobe or much longer, spreading at right angles from axis of lip |
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(5) |
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Lateral lobes of lip oblong, triangular, ovate-triangular, or triangular-lanceolate, shorter than to subequal in length to mid-lobe, spreading at an acute angle to axis of lip |
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(9) |
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| 5 (4) |
Lip lacking a transverse ridge, not clearly demarcated into a hypochile and epichile, deeply 3-lobed near base |
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(6) |
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Lip divided near base by a transverse ridge into a short, concave hypochile and a longer, deflexed, deeply 3-lobed epichile |
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(7) |
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| 6 (5) |
Lateral sepals spreading; lip with a large, subglobose callus near base; spur saccate-ovoid, neck contracted, ca. 2 mm. |
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4 P. formosanus |
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Lateral sepals reflexed; lip ecallose; spur cylindric-clavate, ca. 3.5 mm. |
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5 P. jinchuanicus |
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| 7 (5) |
Lateral lobes of lip linear, 4-7 mm, 2-3 × as long as mid-lobe. |
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3 P. densus |
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Lateral lobes of lip filiform, 8-18 mm, 4-6 × as long as mid-lobe |
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(8) |
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| 8 (7) |
Spur fusiform, 3-5 mm. |
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2 P. calcaratus |
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Spur globose-saccate, 1-2.5 mm. |
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1 P. tentaculatus |
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| 9 (4) |
Lip with a large, subglobose callus near base |
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(10) |
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Lip ecallose |
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(12) |
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| 10 (9) |
Ovary 7-10 mm; dorsal sepal 2.8-3.5 mm; lip 3.5-4.2 mm; spur ovoid, apex acute. |
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14 P. lacertifer |
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Ovary 4-5 mm; dorsal sepal 2-2.5 mm; lip 2-3 mm; spur globose or saccate, apex obtuse |
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(11) |
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| 11 (10) |
Flowers white or pale whitish green; callus on lip glabrous. |
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15 P. coeloceras |
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Flowers green; callus on lip papillose. |
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16 P. neotineoides |
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| 12 (9) |
Plants 1-leaved. |
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6 P. fallax |
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Plants 2-6-leaved |
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(13) |
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| 13 (12) |
Leaves linear, to 0.6 cm wide |
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(14) |
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Leaves elliptic-lanceolate, elliptic, oblong-elliptic, ovate-lanceolate, or ovate, 0.8-6.5 cm wide |
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(15) |
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| 14 (13) |
Petals ca. 3.5 mm; lip 3-4.5 mm; spur cylindric, curved forward, 4-6 mm, almost as long as ovary. |
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7 P. bulleyi |
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Petals 2-2.5 mm; lip 2-2.5 mm; spur scrotiform, 1-1.5 mm, much shorter than ovary. |
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8 P. mannii |
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| 15 (13) |
Petals and lip pure white; dorsal sepal 7-9 mm; lip 9-11 mm. |
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9 P. constrictus |
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Petals and lip green, yellowish green, or creamy white; dorsal sepal 2-5.5 mm; lip 2.5-6 mm |
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(16) |
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| 16 (15) |
Plants usually robust; lip shallowly 3-lobed toward apex, 3.5-4 mm wide, disk with a dark, triangular nectary toward base. |
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10 P. goodyeroides |
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Plants slender to somewhat robust; lip 3-lobed near middle, 1-2 mm wide, disk lacking a nectary |
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(17) |
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| 17 (16) |
Petals and lip creamy white; lip base concave-saccate, papillose in front of spur mouth. |
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11 P. affinis |
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Petals and lip green to yellowish green; lip base shallowly concave, glabrous |
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(18) |
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| 18 (17) |
Leaves narrowly elliptic-lanceolate, 0.8-2 cm wide; floral bracts 5-8 mm, almost as long as ovary; lip ca. 3 mm; spur ca. 1 mm. |
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12 P. elisabethae |
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Leaves elliptic-lanceolate to ovate, 2-3.5 cm wide; floral bracts 8-12 mm, exceeding ovary (and sometimes flowers); lip 4-5.5 mm; spur 1.8-2.5 mm. |
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13 P. parishii |
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List of lower taxa
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