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8. Pertya Schultz Bipontinus, Bonplandia. 10: 109. 1862.
帚菊属 zhou ju shu
Authors: Tiangang Gao & D. J. Nicholas Hind
Macroclinidium Maximowicz.
Shrubs, subshrubs, or perennial herbs, rarely scandent shrubs. Leaves alternate, or tufted on brachyblasts; leaf blade linear, subulate, lanceolate, oblong, or ovate, entire, dentate, or denticulate. Capitula subsessile or pedunculate, in glomerulate, corymbose, or paniculate synflorescences, or solitary, discoid, homogamous; involucre campanulate or cylindric; phyllaries many, 3-seriate to multiseriate, imbricate, unequal, herbaceous or leathery; receptacle flat, alveolate or not, glabrous or villous around, epaleate. Florets few, bisexual, rarely unisexual, tubular, corollas deeply 5-lobed, regularly or slightly irregularly, rarely obviously irregularly, lobed, lobes linear, revolute; anther bases long tailed, apex appendaged; styles shallowly divided at apex, style branches very short, abaxially with short hairs or papillae, apices obtuse. Achenes terete, obovoid, or obconic, 10-ribbed; pappus setae barbellate, numerous. 2n = 24, 26, 28.
About 25 species: Afghanistan, China, Japan, Thailand; 17 species (16 endemic) in China.
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1 |
Leaf blade ovate to broadly ovate, rarely subelliptic on brachyblasts, (2.5-)3-7.5 mm wide, margin serrate, veins 3 |
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(2) |
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Leaf blade linear, oblong, linear-lanceolate, or ovate-lanceolate, rarely subelliptic, 1-6(-15) mm wide, margin entire, vein 1, never 3, rarely with weak lateral veins |
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(9) |
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2 (1) |
Leaves alternate or tufted; capitulum solitary |
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(3) |
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Leaves alternate; capitula many, rarely few, in glomerulate or corymbose synflorescences |
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(4) |
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3 (2) |
Brachyblast leaves on branchlets elliptic; capitula terminal on branchlets. |
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1 P. scandens |
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Brachyblast leaves on branchlets ovate; capitula terminal on long shoots. |
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2 P. simozawae |
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4 (2) |
Capitulum 1-flowered; involucres cylindric, ca. 3 mm in diam. |
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3 P. desmocephala |
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Capitulum 4-12-flowered; involucres campanulate, 5-12 mm in diam |
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(5) |
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5 (4) |
Phyllary apices mucronate to acuminate, pungent. |
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4 P. pungens |
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Phyllary apices rounded to obtuse, never pungent |
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(6) |
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6 (5) |
Capitula solitary, paired, or in corymbose synflorescences; leaf blade base broadly cuneate, obtuse, or truncate |
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(7) |
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Capitula 3-8 in glomerulate synflorescences; leaf blade base cordate, rarely truncate |
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(8) |
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7 (6) |
Florets all bisexual; involucres narrowly campanulate, 5-6 mm in diam.; achenes glabrous or apices sparsely pubescent, pappus dark white. |
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5 P. henanensis |
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Marginal florets female, central florets bisexual; involucres broadly campanulate, ca. 12 mm in diam.; achenes densely hairy, pappus brownish. |
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6 P. corymbosa |
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8 (6) |
Leaf blade apex mucronate or obtuse; florets 9-12 per capitulum. |
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7 P. pubescens |
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Leaf blade apex acuminate to caudate; florets 4 or 5 per capitulum. |
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8 P. cordifolia |
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9 (1) |
Phyllaries multiseriate, at least 6- or 7-seriate; leaf margin strongly revolute |
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(10) |
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Phyllaries 3- or 4-seriate, rarely 5-seriate, few; leaf blade flat |
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(13) |
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10 (9) |
Capitula many and small, 10-15 × 7-10 mm, solitary and terminal on branchlets or subtended by tufted leaves; florets 4-6 per capitulum |
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(11) |
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Capitula few and large, 2.5-3 × 1.5-2.5 cm, solitary and terminal on branches; florets 7-11 per capitulum |
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(12) |
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11 (10) |
Leaves on branchlets homogeneous, leaf margin strongly revolute, terete, apex mucronate. |
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9 P. phylicoides |
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Leaves on branchlets heterogeneous, leaf margin strongly or slightly revolute, dimorphic, one form terete, other form oblong or spatulate-oblong, apex obtuse or rounded. |
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10 P. berberidoides |
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12 (10) |
Capitula terminal, long pedunculate; phyllaries ca. 6-seriate; leaves subterete. |
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11 P. monocephala |
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Capitula terminal or axillary, sessile; phyllaries 16-18-seriate; leaves on branchlets oblong. |
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12 P. tsoongiana |
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13 (9) |
Capitula 1-flowered |
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(14) |
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Capitula multiflowered or with only 2 florets in female plants |
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(15) |
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14 (13) |
Leaf blade oblong to linear, 11-40 × 2-5 mm; capitula solitary, terminal on branchlets; phyllaries 3-seriate. |
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13 P. uniflora |
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Leaf blade elliptic or obovate, 2-12 × 1-6 mm; capitula many, in leafy panicles, axillary or terminal; phyllaries 5-7-seriate. |
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14 P. bodinieri |
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15 (13) |
Involucres narrow, 2-3 mm in diam. |
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15 P. discolor |
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Involucres wide, 5-8 mm in diam |
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(16) |
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16 (15) |
Leaves without lateral veins and reticulate veins. |
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16 P. angustifolia |
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Leaves with obvious lateral veins and reticulate veins. |
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17 P. sinensis |
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List of lower taxa
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