1 |
Plants of high altitudes, (3500–)4000–5000 m, short and squat 5–20(–40) cm; stems usually much reduced, often thickened, unbranched. |
|
(2) |
+ |
Plants from varying altitudes and habitats, usually much larger (0.3–2 m), smaller specimens slender-stemmed. |
|
(10) |
|
|
|
|
2 (1) |
Bracteole apex acute, acuminate or obtuse. |
|
(3) |
+ |
Bracteole apex 1–2-pinnate/pinnatifid. |
|
(5) |
|
|
|
|
3 (2) |
Fruit wings narrow, crisped; calyx teeth lanceolate; rays thickened, particularly at the distal ends; pedicels flattened and winged. |
|
1 Pleurospermum nubigenum |
+ |
Fruit wings narrow or broad, plane; calyx teeth triangular or obsolete; rays uniform thickness, pedicels terete. |
|
(4) |
|
|
|
|
4 (3) |
Petals white; fruit thinly winged; calyx teeth triangular; vittae 3 per furrow. |
|
2 Pleurospermum pulszkyi |
+ |
Petals purple-red; fruit broadly winged; calyx teeth obsolete; vittae 2 per furrow. |
|
3 Pleurospermum lindleyanum |
|
|
|
|
5 (2) |
Bracteole apex 2-pinnate/pinnatifid; ultimate leaf segments linear, 0.3–0.5 mm broad; pedicels flattened. |
|
4 Pleurospermum heterosciadium |
+ |
Bracteole apex 1-pinnate/pinnatifid; ultimate leaf segments lanceolate, 0.5–5 mm broad; pedicels terete. |
|
(6) |
|
|
|
|
6 (5) |
Compact rosette, stemless; inflorescence densely capitate, subsessile; rays reduced, crowded; fruit wings broad and sinuate, not dentate. |
|
5 Pleurospermum hedinii |
+ |
Stems usually distinct; inflorescences not densely capitate, usually distinctly pedunculate; rays not reduced, lax; fruit ribs prominent or irregularly dentate or sinuate winged. |
|
(7) |
|
|
|
|
7 (6) |
Umbels usually subsessile; rays very unequal; fruit tuberculate, wings irregularly dentate. |
|
8 Pleurospermum astrantioideum |
+ |
Umbels distinctly pedunculate; rays subequal; if fruit winged then not tuberculate. |
|
(8) |
|
|
|
|
8 (7) |
Plants with strong unpleasant odor; rays 14–30; fruit wings irregularly dentate. |
|
9 Pleurospermum foetens |
+ |
Plant without unpleasant odor; rays 5–15(–20); fruit ribs prominent to narrowly winged, wing margin entire. |
|
(9) |
|
|
|
|
9 (8) |
Leaves 1–2-pinnate, ultimate segments ovate or orbicular; fruit ribs all broadly winged. |
|
6 Pleurospermum stellatum |
+ |
Leaves 2–3-pinnate, ultimate segments linear-lanceolate; fruit ribs prominent or narrowly sinuate-
winged. |
|
7 Pleurospermum nanum |
|
|
|
|
10 (1) |
Large, often robust plants, (0.5–)0.8–2 m, rarely shorter. |
|
(11) |
+ |
Small, slender plants, 20–40(–50) cm rarely taller. |
|
(23) |
|
|
|
|
11 (10) |
Leaflets of lower leaves broadly ovate, regularly serrate, base truncate or cordate. |
|
(12) |
+ |
Leaflets of lower leaves variously lobed and dissected, often pinnatifid, not broadly ovate, bases usually cuneate to decurrent. |
|
(13) |
|
|
|
|
12 (11) |
Umbels 8–10 cm across; rays 15–20, 4–7 cm; fruit 8–10 × 4–6 mm. |
|
10 Pleurospermum rivulorum |
+ |
Umbels 2–4 cm across; rays 6–8(–15), 0.6–2 cm; fruit 7–8 × 3–4 mm. |
|
11 Pleurospermum rotundatum |
|
|
|
|
13 (11) |
Plants usually very robust, thick-stemmed (except P. stylosum); umbels (7–)12–28 cm across; rays (15–)25–40. |
|
(14) |
+ |
Plants less robust, slender-stemmed; umbels 3–15 cm across; rays 7–28. |
|
(16) |
|
|
|
|
14 (13) |
Rays 15–35; fruit 4–6 mm; ribs sinuate-winged (Xinjiang). |
|
12 Pleurospermum stylosum |
+ |
Rays 20–40(–60); fruit 6–10 mm; ribs cristate- or plane-winged. |
|
(15) |
|
|
|
|
15 (14) |
Ultimate leaf segments pinnatifid, lobes linear; calyx teeth conspicuous, triangular-ovate; fruit wings thick, cristate (NW China). |
|
13 Pleurospermum uralense |
+ |
Ultimate leaf segments coarsely dentate, lobes ovate; calyx teeth obsolete; fruit wings thin, broad
and plane (SW China). |
|
14 Pleurospermum aromaticum |
|
|
|
|
16 (13) |
Petioles of upper leaves expanded into auriculate sheaths; fruit oblong, 8–15 mm. |
|
(17) |
+ |
Petioles of upper leaves expanded or not, but not auriculate; fruit ovate to oblong-ovate, 3.5–8(–10) mm. |
|
(18) |
|
|
|
|
17 (16) |
Leaves irregularly serrate; bracteoles ca. 5 mm broad; seed face deeply concave. |
|
15 Pleurospermum angelicoides |
+ |
Leaves regularly crenate; bracteoles ca. 1 mm broad; seed face slightly concave. |
|
16 Pleurospermum longicarpum |
|
|
|
|
18 (16) |
Leaves 2–4-ternate/pinnate. |
|
(19) |
+ |
Leaves 1–2-ternate/pinnate (3–5-partite to 2-ternate). |
|
(20) |
|
|
|
|
19 (18) |
Leaves with narrow ultimate segments 2–5 mm broad, hirtellous along the margins, lobes round,
obtuse; primary umbel overtopped by laterals; bracteoles oblong to obovate, entire to 3-lobed
at apex, lobes obtuse; petals white; fruit 4–5 mm. |
|
19 Pleurospermum franchetianum |
+ |
Leaves with ultimate segments 0.3–1 mm broad, margin glabrous, lobes lacerate, acute; primary umbel not overtopped by laterals; bracteoles oblanceolate, 3-lobed to pinnate/pinnatifid at apex, apex acute; petals white to pink; fruit 6–10 mm. |
|
20 Pleurospermum benthamii |
|
|
|
|
20 (18) |
Leaves 3–5-partite (sometimes to 2-ternate). |
|
(21) |
+ |
Leaves 2-ternate/pinnate. |
|
(22) |
|
|
|
|
21 (20) |
Leaves 3–5-partite (sometimes to 2-ternate); leaf bases and veins with coarse white hairs reminiscent of those of Heracleum; rays 2–4 cm; petals white. |
|
17 Pleurospermum heracleifolium |
+ |
Leaves always 3-partite, without white hairs; rays 5–6 cm; petals pink. |
|
18 Pleurospermum macrochlaenum |
|
|
|
|
22 (20) |
Rays 10–18, 2–7 cm; bracteoles lanceolate, acute or 3-lobed; fruit smooth, wings sinuate but not irregularly dentate. |
|
21 Pleurospermum decurrens |
+ |
Rays 7–13. 1–2.5 cm; bracteoles linear-lanceolate, entire; fruits tuberculate with irregularly
dentate wings. |
|
22 Pleurospermum cristatum |
|
|
|
|
23 (10) |
Petiole wings greatly expanded, white with purple veining, upper leaves and bracts enveloping the
inflorescence. |
|
23 Pleurospermum amabile |
+ |
Petiole wings not greatly expanded, not purple veined. |
|
(24) |
|
|
|
|
24 (23) |
Leaves 1-pinnate to 2-pinnatifid; rays 2–4; bracteoles broadly ovate, entire. |
|
24 Pleurospermum bicolor |
+ |
Leaves 2–4-ternate/pinnate; rays (4–)6–25; bracteoles entire or more usually pinnatifid at apex. |
|
(25) |
|
|
|
|
25 (24) |
Stems, petioles and leaf rachis conspicuously white pubescent. |
|
26 Pleurospermum pilosum |
+ |
Plants essentially glabrous, bases of umbels and/or petioles sometimes puberulent. |
|
(26) |
|
|
|
|
26 (25) |
Calyx teeth conspicuous, triangular to lanceolate. |
|
(27) |
+ |
Calyx teeth obsolete or minute. |
|
(34) |
|
|
|
|
27 (26) |
Ultimate leaf segments 5–20 mm broad, irregularly serrate or incised. |
|
25 Pleurospermum yunnanense |
+ |
Ultimate leaf segments finely dissected, 1–8(–10) mm broad. |
|
(28) |
|
|
|
|
28 (27) |
Calyx teeth linear-lanceolate; sheaths of upper leaves and bracts broadly auriculate; bracteoles
silvery white with a narrow green midrib, apex acuminate to 3-lobed. |
|
27 Pleurospermum album |
+ |
Calyx teeth triangular or rounded; leaf sheaths somewhat expanded but not auriculate; bracteoles usually white margined, but not conspicuously silvery white, apex usually pinnatifid. |
|
(29) |
|
|
|
|
29 (28) |
Rays 4–7, very unequal; bracteoles entire, apex acuminate. |
|
29 Pleurospermum rupestre |
+ |
Rays (6–)8–20(–30), subequal (but see P. szechenyii); bracteole apex pinnatifid (rarely acuminate). |
|
(30) |
|
|
|
|
30 (29) |
Rays (15–)20–30. |
|
(31) |
+ |
Rays 6–12(–15). |
|
(32) |
|
|
|
|
31 (30) |
Plant without unpleasant odor when crushed; rays very unequal; fruit wings broad, sinuate but not dentate. |
|
30 Pleurospermum szechenyii |
+ |
Plant with a strong unpleasant odor when crushed; rays subequal; fruit wings broad, irregularly dentate. |
|
9 Pleurospermum foetens |
|
|
|
|
32 (30) |
Leaf blades oblong-lanceolate in outline, approaching 2–3-ternate/pinnate; fruit wings broad, sinuate. |
|
31 Pleurospermum wilsonii |
+ |
Leaf blades broadly ovate in outline, 3–4-ternate/pinnate; fruit ribs prominent to winged,
but not sinuate. |
|
(33) |
|
|
|
|
33 (32) |
Ultimate leaf segments ca. 2 mm; calyx teeth ca. 1 mm, green. |
|
32 Pleurospermum hookeri |
+ |
Ultimate leaf segments 3–5 mm; calyx teeth ca. 0.3 mm, dark purple. |
|
33 Pleurospermum tsekuense |
|
|
|
|
34 (26) |
Rays 5–9(–11); bracts and bracteoles entire, acuminate. |
|
(35) |
+ |
Rays (9–)11–25; bracts and bracteoles pinnate/pinnatifid at apex, rarely acuminate. |
|
(36) |
|
|
|
|
35 (34) |
Stems and leaf sheaths purple-red, rays subequal; bracteoles broad, longer than the flowers. |
|
35 Pleurospermum apiolens |
+ |
Stems and leaf sheaths green, rays very unequal; bracteoles linear-lanceolate, shorter than the
pedicels. |
|
28 Pleurospermum simplex |
|
|
|
|
36 (34) |
Larger plants, 30–60 cm; rays 10–25; fruit wings broad, sinuate or irregularly cristate/dentate. |
|
(37) |
+ |
Small slender plants, 20–35(–45) cm; rays 9–15; fruit ribs prominent to narrowly winged, but not sinuate or dentate. |
|
(38) |
|
|
|
|
37 (36) |
Rays subequal; fruit wings sinuate. |
|
37 Pleurospermum linearilobum |
+ |
Rays very unequal; fruit wings irregularly cristate/dentate. |
|
39 Pleurospermum wrightianum |
|
|
|
|
38 (36) |
Rays 1.5–2.5 cm. |
|
34 Pleurospermum giraldii |
+ |
Rays 4–12 cm. |
|
(39) |
|
|
|
|
39 (38) |
Stem branches often opposite or whorled; leaf rachis glabrous; ovary smooth. |
|
36 Pleurospermum handelii |
+ |
Stem branches usually alternate; leaf rachis tuberculate on abaxial surface; ovary tuberculate. |
|
38 Pleurospermum calcareum |