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FOC | Family List | FOC Vol. 12 | Theaceae

2. Pyrenaria Blume, Bijdr. 1119. 1826.

核果茶属 he guo cha shu

Parapyrenaria Hung T. Chang; Sinopyrenaria Hu; Tutcheria Dunn.

Shrubs or trees, evergreen. Leaves petiolate; leaf blade margin serrate. Flowers axillary, solitary, pedicellate or subsessile. Bracteoles 2 or rarely more, caducous or persistent, sometimes clearly differentiated from sepals. Sepals 5(or 6), imbricate, persistent or caducous, unequal in size, outside pubescent to velutinous, inside brown and glabrous. Petals 5 or rarely more, white or rarely pale yellow, basally slightly connate and adnate to stamens. Stamens numerous, glabrous; outer filament whorl basally slightly connate; anthers dorsifixed, 2-loculed, longitudinally cleft. Ovary superior, (2 or)3-5(or 6)-loculed with 2-5 ovules per locule, placentation axile, tomentose; styles 3-5, distinct or basally connate. Fruit a loculicidal capsule dehiscent from base or drupaceous and indehiscent, usually 2 seeds per locule; pericarp woody or fleshy; columella persistent. Seeds longitudinally compressed, wingless; testa bony, smooth, shiny; hilum linear; endosperm absent; embryo large; cotyledons thin.

About 26 species: China, NE India, Indonesia, Japan (Ryukyu Islands), Malaysia, Myanmar, Philippines, Thailand, Vietnam; 13 species (seven endemic) in China.

1 Ovary 5-loculed; styles 5, distinct; fruit drupaceous, indehiscent, apically 5-lobed   (2)
+ Ovary 3-5(or 6)-loculed; styles connate; fruit a capsule or drupaceous and indehiscent, apically unlobed   (5)
2 (1) Flowers subsessile; bracteoles and sepals more than 40, not clearly differentiated.   4 P. oblongicarpa
+ Flowers pedicellate; bracteoles 2 and sepals 5 or 6, clearly differentiated   (3)
3 (2) Leaf blade 15-33 × 7-14 cm, base rounded; bracteoles and sepals not leaflike; flowers ca. 7 cm in diam.; fruit globose, 5-8 cm in diam.   1 P. menglaensis
+ Leaf blade 8-16 × 3-6 cm, base cuneate; bracteoles and sepals leaflike; flowers 2-4 cm in diam.; fruit oblate, subglobose, or long obovoid, less than 4 cm in diam   (4)
4 (3) Leaf blade oblong to obovate-oblong, papery, yellowish green when dry, secondary veins raised on both surfaces; fruit long obovoid.   2 P. diospyricarpa
+ Leaf blade oblanceolate to oblong-lanceolate, leathery, adaxially yellowish brown when dry, secondary veins adaxially slightly impressed; fruit oblate to subglobose.   3 P. khasiana
5 (1) Flowers subsessile (pedicel 3-6 mm in P. kwangsiensis); bracteoles and sepals not clearly differentiated; fruit indehiscent and drupaceous   (6)
+ Flowers distinctly pedicellate; bracteoles and sepals clearly differentiated; fruit a dehiscent capsule   (7)
6 (5) Sepals 5; petals 5; ovary and fruit (4 or)5-loculed.   12 P. kwangsiensis
+ Sepals 8-10; petals 8-10; ovary and fruit 3-loculed.   13 P. jonquieriana subsp. multisepala
7 (5) Fruit 1-2 cm in diam   (8)
+ Fruit 2-8 cm in diam   (10)
8 (7) Flowers 5-6 cm in diam.; pedicel 10-15 mm.   6 P. wuana
+ Flowers 2-4 cm in diam.; pedicel 1-4(-8) mm   (9)
9 (8) Flowers ca. 4 cm in diam.; year-old branchlets hirsute; leaf blade adaxially hirtellous along midvein; sepals outside hirsute, apex acute; stamens ca. 10 mm; style hirsute.   9 P. hirta
+ Flowers 2-2.5 cm in diam.; year-old branchlets glabrescent; leaf blade adaxially glabrous; sepals outside sericeous, apex rounded; stamens 6-8 mm; style glabrous.   10 P. microcarpa
10 (7) Current year branchlets pubescent or villous   (11)
+ Current year branchlets glabrous   (12)
11 (10) Leaf blade 7-12.5 cm; bracteoles alternate, leaflike, oblong, ca. 2.5 cm; sepals 5, broadly ovate, 1-1.5 cm.   5 P. sophiae
+ Leaf blade 16-18 cm; bracteoles opposite, not leaflike, ovate to broadly ovate, 1-1.5 cm; sepals 9-11, suborbicular, 2-3(-4) cm.   7 P. spectabilis
12 (10) Current year branchlets rust colored, not maculate; capsule apex convex.   8 P. pingpienensis
+ Current year branchlets grayish white and dark purplish brown maculate; capsule apex abruptly pointed.   11 P. maculatoclada

  • List of lower taxa


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