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FOC | Family List | FOC Vol. 12 | Rhamnaceae

7. Sageretia Brongniart, Mém. Fam. Rhamnées. 52. 1826.

雀梅藤属 que mei teng shu

Ampeloplis Rafinesque.

Shrubs scandent or erect, rarely small trees, unarmed or spinescent. Branchlets alternate or subopposite, often terminating in a woody spine. Leaves alternate or subopposite; stipules small, caducous; leaf blade papery to leathery, pinnately veined, margin serrate, rarely entire. Flowers mostly very small, 1-2 mm in diam., bisexual, 5-merous, usually sessile or subsessile, rarely pedicellate, in spikes or spicate panicles, rarely in racemes. Calyx tube shallowly cup-shaped to hemispherical; sepals triangular, ± fleshy, adaxially medially keeled and hooded. Petals spatulate, apex 2-lobed to ± deeply emarginate. Stamens equaling petals or slightly longer; anthers dorsifixed. Disk cup-shaped, thick, fleshy, outer margin free from calyx tube, ± distinctly erect, entire or 5-lobed. Ovary superior, 2- or 3-loculed, with 1 ovule per locule; style short, stout, undivided, apically ± distinctly 2- or 3-lobed. Drupe obovoid-globose, with 2 or 3 one-seeded stones, base with remnants of persistent calyx tube. Seeds compressed, slightly asymmetrical, concave at both ends.

About 35 species: mainly in SE Asia, a few species in Africa and North America; 19 species (15 endemic) in China.

Sageretia gongshanensis G. S. Fan & L. L. Deng (Sida 17: 677. 1997) and S. latifolia Handel-Mazzetti (Symb. Sin. 7: 673. 1933), both described from Yunnan, and S. yunlongensis G. S. Fan & L. L. Deng (Sida 16: 477. 1995), described from Xizang and Yunnan, could not be treated here because no material was seen by the authors.

The fruit of some species is edible. The leaves are used as a substitute for tea. Several species are very popular in Bonsai gardening.


1 Flowers distinctly pedicellate, in racemes or panicles; fruit maturity delayed to following year   (2)
+ Flowers sessile or subsessile, in spikes or spicate panicles; fruit maturity in current year   (3)
       
2 (1) Leaves 5-12 × 2.5-5 cm, with 5 or 6(or 7) pairs of lateral veins.   18 S. henryi
+ Leaves distinctly smaller, 0.8-2 × 0.7-1.2 cm, with 3 or 4 pairs of lateral veins.   19 S. pedicellata
       
3 (1) Rachis of inflorescence glabrous, rarely sparsely pubescent   (4)
+ Rachis of inflorescence tomentose or densely pubescent   (9)
       
4 (3) Leaf blade 0.5-2(-2.5) cm × less than 1.4 cm, petiole 1-3 mm, lateral veins 3-5 pairs   (5)
+ Leaf blade larger than 2.5 × 1.4 cm, petiole longer than 4 mm, lateral veins 2-8 pairs   (6)
       
5 (4) Leaves papery, often fascicled, leaf blade obovate or oblong, apex emarginate or rounded, margin entire or crenate; inflorescences usually inserted at middle of branches.   1 S. horrida
+ Leaves leathery, not fascicled, but often distichous, leaf blade oblong or ovate-elliptic, apex obtuse or acute, not emarginate, margin minutely serrulate; inflorescences at ends of branchlets.   2 S. pycnophylla
       
6 (4) Leaves adaxially smooth; lateral veins few, 2 or 3(or 4) pairs.   3 S. paucicostata
+ Leaves adaxially shiny; lateral veins 5-8 pairs   (7)
       
7 (6) Leaves abaxially barbate at vein axils, leaf bases asymmetric.   4 S. lucida
+ Leaves abaxially glabrous, leaf bases symmetric   (8)
       
8 (7) Leaves papery or subleathery, apex acuminate or acute, margin not revolute; flowers usually in terminal spicate-paniculate inflorescences; fruit maturity in following year.   5 S. gracilis
+ Leaves leathery or thickly so, apex obtuse or rounded, margin often revolute; flowers in axillary or terminal lax spikes; fruit maturity within current year.   6 S. camelliifolia
       
9 (3) Leaf blade abaxially glabrous or pubescent on veins, or barbate at vein axils   (10)
+ Leaf blade abaxially tomentose, hairs persistent or ± deciduous   (14)
       
10 (9) Leaves up to 4.5 × 2.5 cm, lateral veins 3-5(-7) pairs, adaxially not impressed.   13 S. thea
+ Leaves 4-15 cm × more than 3.5 cm, lateral veins 5-10 pairs, adaxially distinctly impressed   (11)
       
11 (10) Branchlets often with hooked, incurved long spines; leaf blade usually oblong, abaxially barbate at axils of veins, petioles glabrous; fruit ripe in current year   (12)
+ Branchlets with erect spines or without spines; leaf blade usually ovate-elliptic, glabrous or abaxially pubescent on veins, petioles pubescent or sparsely pubescent; fruit ripe in following year   (13)
       
12 (11) Leaves leathery, major veins adaxially distinctly impressed.   14 S. hamosa
+ Leaves papery, major veins adaxially inconspicuously impressed.   15 S. randaiensis
       
13 (11) Leaf blade leathery, abaxially glabrous, lateral veins 5-7 pairs, apex acuminate, rarely acute, base subrounded, slightly asymmetric.   16 S. melliana
+ Leaf blade papery or thinly leathery, abaxially on veins pubescent, lateral veins 7-10 pairs, apex caudate-acuminate or long acuminate, base cordate or subrounded, never asymmetric.   17 S. subcaudata
       
14 (9) Leaf blade 1-2.5 × 0.3-1.5 cm, abaxially persistently densely tomentose   (15)
+ Leaf blade larger than 3 × 1 cm, often ferruginous or canescent tomentose, hairs deciduous or not   (17)
       
15 (14) Leaf blade papery, margin explanate, lateral veins 4-6 pairs.   7 S. brandrethiana
+ Leaf blade leathery, margin revolute or not, lateral veins 3 or 4 pairs   (16)
       
16 (15) Leaf blade dimorphic, large one 1.5-2.5 × 0.8-1.5 cm, small one 1-1.5 × 0.3-0.6 cm, all thickly leathery, margin revolute; calyx and ovary tomentose.   8 S. lijiangensis
+ Leaf blade uniform, 1-1.5 × 0.7-1 cm, thinly leathery, margin explanate; calyx puberulous; ovary glabrous.   9 S. yilinii
       
17 (14) Leaf blade oblong, ovate-oblong, or ovate, apex acute, adaxially distinctly rugose.   10 S. rugosa
+ Leaf blade lanceolate, lanceolate-oblong, or lanceolate-elliptic, apex acuminate or long acuminate, adaxially not rugose   (18)
       
18 (17) Leaves abaxially incanous or ferruginous tomentose, lateral veins 7-9 pairs; branchlets and petioles incanous or ferruginous tomentose.   11 S. omeiensis
+ Leaves abaxially ferruginous tomentose, hairs later partly or wholly deciduous, lateral veins 5 or 6 pairs; branchlets and petioles ferruginous pubescent.   12 S. laxiflora

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