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FOC | Family List | FOC Vol. 9 | Rosaceae

45. SANGUISORBA Linnaeus, Sp. Pl. 1: 116. 1753.

地榆属 di yu shu

Li Chaoluan (Li Chao-luang); Hiroshi Ikeda, Hideaki Ohba

Sanguisorba filiformis

Credit: Harvard University Herbaria

Poterium Linnaeus.

Herbs perennial. Rootstock robust, bearing many fusiform, cylindric roots in lower part. Stipules sheathing, adnate to petiole, petiole sheathing and imbricate at base; leaf blade imparipinnate; leaflets serrate at margin. Inflorescences terminal on elongate scapes, densely capitate or spicate, bracteate and bracteolate. Flowers bisexual, rarely unisexual (when plants monoecious). Hypanthium with a constricted throat. Sepals 4(–7), imbricate, petaloid, purple, red, pink, or white, rarely greenish. Petals absent. Disk lining hypanthium. Stamens usually 4, rarely more, inserted in throat of hypanthium; filaments free, rarely partly connate; anthers didymous. Carpel 1(or 2), included in hypanthium; ovule pendulous; style terminal, filiform; stigma penicillate. Achene dry, included in hardened, muricate or winged hypanthium. Cotyledons planoconvex. x = 7.

About 30 species: Asia, Europe, North America; seven species (one endemic) in China.

1 Inflorescences flowering from base upward.   (2)
+ Inflorescences flowering from apex downward.   (3)
2 (1) Leaflets of radical leaves usually truncate to subcordate at base; inflorescences usually nodding, thick; sepals white or pinkish; bracts longer than hypanthium in bud, densely hairy; filaments gradually dilated from base, broadest near middle, then gradually attenuate toward apex, narrower than anther, or sometimes little dilated.   6 S. alpina
+ Leaflets of radical leaves cordate or deeply so at base; inflorescences usually erect, long, slender; sepals white; bracts nearly equaling hypanthium in bud, abaxially sparsely hairy or subglabrous, margin ciliate; filaments compressed-dilated from middle to apex, broadest at apex, nearly as broad as anther.   7 S. stipulata
3 (1) Inflorescences capitate; sepals white.   (4)
+ Inflorescences ellipsoid, cylindric, or elongate; sepals purple, red, pink or white.   (5)
4 (3) Plants less than 35 cm tall; leaflets of radical leaves broadly ovate to suborbicular; flowers unisexual (females in center, males surrounding); style 0.5–1 × as long as sepals; fruiting hypanthium with 4 ribs.   4 S. filiformis
+ Plants to 85 cm tall; leaflets of radical leaves ovate, elliptic, or long elliptic; flowers bisexual; style equaling sepals; fruiting hypanthium with 4 winged ribs.   5 S. diandra
5 (3) Filaments filiform, subequaling sepals.   1 S. officinalis
+ Filaments compressed-dilated, 0.5–2 × as long as sepals.   (6)
6 (5) Leaflets of radical leaves linear-lanceolate, base subcordate or rounded to obliquely broadly cuneate, margin acutely incised serrate.   2 S. tenuifolia
+ Leaflets of radical leaves ovate, elliptical, or oblong, base cordate, margin coarsely obtusely serrate.   3 S. applanata

  • List of lower taxa


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