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5. Streblus Loureiro, Fl. Cochinch. 2: 754. 1790.
鹊肾树属 que shen shu shu
Dimerocarpus Gagnepain; Diplocos Bureau; Diplothorax Gagnepain; Epicarpurus Blume; Paratrophis Blume; Pseudostreblus Bureau; Pseudotrophis Warburg; Taxotrophis Blume; Teonongia Stapf.
Trees or shrubs, occasionally scandent shrubs, with latex, sometimes spiny; monoecious or dioecious. Spines when present axillary. Stipules free or connate, lateral to ± amplexicaul, caducous. Leaves alternate, distichous; petiole short; leaf blade with or without cystoliths on one or both surfaces, margin entire or toothed and not glandular; secondary veins pinnate. Male inflorescences axillary, sometimes bisexual with a single central female flower, racemose, spicate, or ± capitate, pedunculate. Female inflorescences often 1-flowered. Male flowers: calyx lobes (3 or)4(or 5), free or basally adnate with pistillode, imbricate or valvate; stamens inflexed in bud; anthers small, reniform, extrorse. Female flowers: calyx lobes 4, opposite, free or ± connate, imbricate; ovary superior, ovoid to oblique, basally thick and fleshy, enclosed by persistent calyx lobes or not; style 2-branched. Fruit free, not aggregated into a syncarp. Drupes globose, dehiscent or indehiscent, sometimes loosely covered by slightly enlarged persistent calyx, often basally fleshy on one side. Seed large, globose, enveloped by membranous endocarp, endosperm present or absent; cotyledons unequal.
Twenty-two species: tropical and subtropical Asia; seven species in China.
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| 1 |
Male inflorescences capitate to shortly racemose; female inflorescences 1-flowered; bisexual inflorescences sometimes present. |
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(2) |
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Male inflorescences spicate or scorpioid cymose; female inflorescences usually 2-8-flowered (1-flowered in S. indicus); bisexual inflorescences usually absent (present in S. indicus). |
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(4) |
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| 2 (1) |
Plants with spines; branchlets with dark-yellow marginal hairs on one side; drupes basally fleshy; plants dioecious. |
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3 S. taxoides |
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Plants spineless; branchlets uniformly hairy; drupes basally not fleshy; plants often monoecious. |
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(3) |
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| 3 (2) |
Leaf blade papery; pistillode cubic; drupe dehiscent, not enclosed by persistent calyx lobes. |
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2 S. tonkinensis |
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Leaf blade leathery; pistillode conic to cylindric; drupes indehiscent, enclosed by persistent calyx lobes. |
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1 S. asper |
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| 4 (1) |
Male inflorescences scorpioid; male flowers 5-merous; female inflorescences 1-flowered; drupes indehiscent. |
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4 S. indicus |
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Male inflorescences racemose or spicate; male flowers 4-merous; female inflorescences 2-8-flowered; drupes ± dehiscent. |
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(5) |
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| 5 (4) |
Leaf blade ovate-oblong, narrowly elliptic, or lanceolate, apex shortly attenuate and tipped by 1 long tooth flanked by 2 short triangular teeth; cotyledons ± equal. |
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5 S. zeylanicus |
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Leaf blade elliptic, obovate-elliptic, rhombic, or oblong-obovate, apex acute, blunt, or retuse, with two spiny teeth or caudate; cotyledons unequal. |
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(6) |
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| 6 (5) |
Branchlets spineless; leaf blade elliptic to obovate-elliptic, 8-16 × 3-6 cm, membranous to papery, margin entire or ± undulate, apex caudate. |
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6 S. macrophyllus |
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Branchlets with 1-1.5(-4.5) cm spines; leaf blade rhombic to oblong-obovate, 1-4.5(-9) × 0.6-2.5(-5) cm, leathery, margin loosely revolute with 5 or fewer spiny teeth, apex acute, blunt, or retuse, with two spiny teeth. |
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7 S. ilicifolius |
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Lower Taxa
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