1 |
Trees, shrubs, or climbers; florets (1 or)2-12 per involucre, rarely more and then plant a climber with axillary synflorescences |
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(2) |
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Erect annual or perennial herbs; florets 19 or more per involucre; synflorescences terminal |
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(16) |
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2 (1) |
Trees or self-supporting shrubs |
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(3) |
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Scandent shrubs |
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(9) |
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3 (2) |
Leaves obovate to oblanceolate, clearly widest above middle, basal secondary veins substraight toward leaf margins; petiole short and broad or expanded into a sheath at base; synflorescence conically thyrsoid (3. Monosis group) |
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(4) |
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Leaves ovate to elliptic or oblong, widest at or below middle, basal secondary veins curved and often becoming subparallel to basal leaf margins; petiole slender; synflorescence flat-topped to rounded |
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(5) |
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4 (3) |
Trees 3-8 m tall; leaf blade to 50 × 20 cm, abaxially glabrous or sparsely pubescent, lateral veins 12-17-paired; achenes 3-4 mm. |
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8 V. volkameriifolia |
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Shrubs 2-3 m tall; leaf blade 15-30 × 4-9 cm, abaxially densely pubescent to fulvous tomentose, lateral veins 7-12-paired; body of mature achenes ca. 2.5 mm. |
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9 V. parishii |
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5 (3) |
Corolla 8-10 mm, uniformly slender from base before widening into limb; achenes strongly 10-ribbed; style base without node; sweeping hairs of style branches pointed (4. Gymnanthemum group). |
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10 V. extensa |
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Corollas 5-7 mm, widening gradually from near base; achenes 5-ribbed to weakly 10-ribbed; style base with large distinct node; sweeping hairs of style branches blunt (1. Strobocalyx group p.p.) |
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(6) |
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6 (5) |
Trees; branches fulvous or brown tomentose; inner phyllaries persistent |
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(7) |
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Shrubs, rarely small trees; branches gray or gray-brown tomentose; innermost phyllaries ± deciduous |
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(8) |
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7 (6) |
Florets 5 or 6 per capitulum; involucres 6-8 mm in diam.; phyllaries ovate-oblong, shortly pubescent or subglabrous; achenes subtriangular in section, 8-10-ribbed, glabrous, glandular; leaf blade thinly leathery, glabrous or sparsely puberulent along veins, margin entire. |
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1 V. arborea |
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Florets ca. 10 per capitulum; involucres 4-5 mm in diam.; phyllaries ovate, abaxially densely tomentose; achenes 4- or 5-angled, glabrous; leaf blade papery, abaxially hispidulous, adaxially tomentose, margin repand or laxly dentate. |
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2 V. solanifolia |
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8 (6) |
Capitula 6-8 mm in diam., florets 8-12; involucres subglobose or hemispheric with rounded base, 4-6 mm in diam.; phyllaries obtuse or rounded at apex, white sericeous; achenes 3.5-4 mm, 3-ribbed ventrally, glabrous; leaves membranous or thinly papery. |
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3 V. bockiana |
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Capitula 2-3 mm in diam., florets 5 or 6; involucres obconical with cuneate base, 2-3 mm in diam.; phyllaries few, acute or slightly acute at apex, sparsely shortly pubescent; achenes ca. 3 mm, slightly ribbed, sparsely puberulent, glandular; leaves hardly papery. |
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4 V. esculenta |
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9 (2) |
Synflorescences terminal and subterminal, forming broad panicle; pappus white or dirty white; branches densely reddish- or yellowish brown pubescent (1. Strobocalyx group p.p.) |
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(10) |
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Synflorescences mostly axillary, forming long thyrse; pappus reddish, yellow, or brown; branches grayish brown pubescent |
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(11) |
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10 (9) |
Branches densely reddish brown pubescent, eglandular; lateral veins 7- or 8-paired; capitula 7-12 mm in diam., with ca. 10 florets, peduncle up to 15 mm; achenes 10-ribbed, eglandular. |
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5 V. sylvatica |
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Branches densely yellowish brown pubescent, glandular; lateral veins 3-6-paired; capitula 3-5 mm in diam., with 2 or 3 florets, peduncle less than 2 mm; achenes 5-ribbed, glandular. |
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6 V. chunii |
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11 (9) |
Hairs of stems and leaves T-shaped with long arms, shortly stalked and pressed against stems or leaf surface, silky; corollas without long slender bases; style with distinct basal node; achenes 5-angled (2. Tarlmounia group). |
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7 V. elliptica |
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Hairs of stems and leaves simple, not T-shaped; corollas with long and filiform bases; bases of styles without nodes; achenes strongly 10-ribbed (5. Decaneuropsis group) |
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(12) |
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12 (11) |
Capitula in terminal or subterminal panicles; receptacle ferruginous pubescent; pappus reddish or reddish brown; leaves papery; branches reddish brown or grayish brown tomentose. |
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11 V. cumingiana |
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Capitula mostly axillary or on short lateral branches, solitary or in groups of 3-5, forming leafy thyrse; receptacle glabrous; pappus yellowish brown; leaves leathery or thinly leathery; branches sparsely, shortly pubescent or glabrescent |
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(13) |
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13 (12) |
Leaf blade oblong or oblong-lanceolate, lateral veins 7-9-paired, adaxially densely gray-green tomentose; capitula with ca. 10 florets; achenes 3-3.5 mm |
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(14) |
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Leaf blade ovate-oblong or oblong-elliptic, lateral veins 5-7-paired, adaxially sparsely pubescent; capitula with 18 or more florets; achenes 4-4.5 mm |
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(15) |
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14 (13) |
Phyllaries linear or linear-lanceolate, acute at apex, with dense short pubescence on margins and tips. |
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12 V. gratiosa |
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Phyllaries oblong, with short acute tips, glabrous. |
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V. andersonii (see note under V. cumingiana)
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15 (13) |
Capitula 7-8 mm in diam., 3-5 in small axillary panicles; florets 20-25, reddish purple; achenes 3-3.5 mm, 10-ribbed, with short patent hairs and glands; leaves thinly leathery, elliptic or ovate-elliptic, entire. |
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13 V. blanda |
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Capitula ca. 20 mm in diam., solitary in leaf axils, racemose; florets many, white; achenes 7-9 mm, 7- or 8-ribbed, glabrous; leaves leathery, obovate-oblong or elliptic-oblong, margin often revolute, sparsely denticulate at upper margins. |
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14 V. chingiana |
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16 (1) |
Achenes 4- or 5-angled to almost terete; phyllary apex acute or aristate-acuminate; plants often annual herbs (7. Cyanthillium group) |
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(17) |
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Achenes (7-)10-ribbed or striate |
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(20) |
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17 (16) |
Achenes unribbed or rarely inconspicuously ribbed, ± compressed, densely white pilose; pappus 2-seriate, outer setae many, short, and persistent; capitula small, ca. 6 mm in diam., many in relatively broad panicles; florets 19-28 |
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(18) |
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Achenes 4- or 5-angled, glabrous and glandular; pappus 1-seriate, deciduous; capitula larger, 6-10 mm in diam., few or 2 or 3 terminal on branches; florets more than 30 |
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(19) |
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18 (17) |
Corolla ca. as long as pappus; leaf blade rhombic to ovate. |
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26 V. cinerea |
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Corolla much longer than pappus at anthesis; leaf blade linear-lanceolate to lanceolate. |
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27 V. benguetensis |
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19 (17) |
Capitula several, laxly corymbose, 5-7 mm in diam.; stem leaves crowded, spatulate, apex rounded or obtuse, margin subentire, abaxially densely white hispidulous; small subshrubs, ca. 10 cm tall. |
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28 V. maritima |
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Capitula usually 2 or 3 terminal on branches or paired, 8-10 mm in diam.; stem leaves widely separated, ovate, ovate-elliptic, rarely orbicular, apex obtuse, slightly acute, margin crenate-shallowly dentate, abaxially sparsely shortly hairy or glabrous, adaxially gray sericeous-pubescent; robust annual herbs, to 1 m tall. |
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29 V. patula |
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20 (16) |
Capitula subtended by foliose bracts as long as phyllaries; phyllary apex ± leaflike; corolla with slender base more than 2 × as long as abruptly wider narrowly campanulate limb; pappus bristles flattened on outer surface (9. Baccharoides group). |
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31 V. anthelmintica |
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Capitula without foliose or membranous appendaged bracts at base of involucre; phyllary apex not leaflike; corolla with slender base up to as long as narrowly funnel-shaped limb; pappus segments rounded and/or scabrid on outer surface |
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(21) |
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21 (20) |
Lower part of leaf blade with mostly spreading secondary veins; slender base of corolla tube ca. as long as limb; base of style without node (8. Khasianthus group). |
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30 V. subsessilis |
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Lower part of leaf blade with secondary veins ascending at 45° angles or more; slender base of corolla tube shorter than limb; base of style with distinct node (6. Acilepis group) |
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(22) |
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22 (21) |
Capitula ca. 30 mm in diam.; phyllaries abaxially pale yellowish white tomentose. |
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25 V. albosquama |
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Capitula 5-25 mm in diam.; phyllaries glabrous to densely pubescent, green, often with darker apex |
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(23) |
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23 (22) |
Outer pappus bristles many, short |
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(24) |
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Outer pappus bristles few to none |
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(27) |
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24 (23) |
Capitula usually more than 5 in terminal, often somewhat elongated, panicle or thyrse, 5-8 mm in diam.; achenes glabrous; pappus white or dirty white. |
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15 V. saligna |
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Capitula few, usually 2 or 3(-5) in ± flat-topped cyme, rarely solitary in leaf axils, 8-15 mm in diam.; achenes hairy and glandular; pappus sordid white or rubescent, outer bristles deciduous or persistent |
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(25) |
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25 (24) |
Stems hispidulous; capitula 10-15 mm in diam., peduncle very short to almost absent; florets ca. 30; phyllary apex apiculate; leaves oblanceolate or obovate-elliptic, sharply serrate, hispidulous on both surfaces. |
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16 V. aspera |
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Stems shortly pubescent or adpressed hairy; capitula 6-8(-10) mm in diam., peduncle 7-15 mm; florets 15-20; phyllary apex acute or acuminate; leaves oblong, oblong-lanceolate, obovate, or obovate-elliptic, denticulate or sparsely serrate toward apex, subglabrous or shortly pubescent along veins only |
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(26) |
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26 (25) |
Involucres hemispheric; phyllaries with apex acute, upper part arachnoid pubescent and glandular; achenes hispidulous; leaves papery, laxly on stems, oblong or oblong-lanceolate, sparsely serrate on upper margins. |
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17 V. clivorum |
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Involucres obconical; phyllaries with apex acuminate, subscarious, margin pubescent; achenes patently villous; leaves leathery, often conferted below flowering branches, obovate or obovate-elliptic, upper leaves linear or linear-lanceolate, denticulate. |
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18 V. attenuata |
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27 (23) |
Capitula 4-6 mm in diam.; florets 8-12; shrubs or shrubby herbs 1-3 m tall; phyllary apex obtuse or bluntly acute, adpressed to form smooth involucre. |
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19 V. divergens |
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Capitula 12-25 mm in diam.; florets many; herbs 15-80(-200) cm tall; phyllary apex acute to acuminate, often conspicuous or colored spinescent, often recurved |
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(28) |
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28 (27) |
Capitula sessile or nearly so, solitary, often axillary and forming spikelike synflorescence; outermost phyllaries almost subulate. |
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24 V. squarrosa |
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Capitula mostly clearly pedunculate, often in flat-topped cymes, never axillary; outer phyllaries lanceolate to ovate |
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(29) |
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29 (28) |
Achenes glabrous; [inner phyllaries ca. 2.5 mm wide; stem hairs pigmented, clearly multicellular] |
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(30) |
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Achenes adpressed hairy; inner phyllaries 1.5-2.2 mm wide; stem hairs pale, slender, obscurely multicellular |
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(31) |
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30 (29) |
Panicles relatively broad, ± flat-topped; phyllaries often recurved at tip. |
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21 V. silhetensis |
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Panicles relatively narrow, ± cylindric; phyllaries straight to slightly incurved. |
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22 V. doichangensis |
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31 (29) |
Involucre 12-15 mm wide; leaf blade thinly papery; achenes eglandular. |
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20 V. nantcianensis |
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Involucre 15-20 mm wide; leaf blade more stiffly papery; achenes glandular. |
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23 V. spirei |
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