1. Protowoodsia manchuriensis (Hooker) Ching, Lingnan Sci. J. 21: 37. 1945.
膀胱蕨 pang guang jue
Woodsia manchuriensis Hooker, Sec. Cent. Ferns, t. 98. 1861; Diacalpe manchuriensis (Hooker) Trevisan; Physematium manchuriense (Hooker) Nakai.
Plants (8-)15-20 cm. Rhizomes short, erect; scales brown, lustrous, ovate-lanceolate or lanceolate, entire. Fronds clustered; stipe brownish stramineous, 2-2.5 cm, less than 1 mm in diam., brittle, usually breaking, sparsely short glandular hairy, with few scales in lower part; lamina 1-pinnate-pinnatifid, lanceolate, 12-18 × 1.5-4 cm, herbaceous, sparsely glandular hairy, base attenuate, apex acuminate; pinnae (12-)16-20 pairs, oblique, subsessile; middle pinnae largest, ovate-lanceolate, 1-1.5 × up to 1 cm, pinnatifid, apex obtuse; lower pinnae shorter than above, widely separate; basal pair of pinnae ovate or flabellate; ultimate segments oblong, undulate or dentate. Veins free, pinnate, not reaching laminar margin. Sori consisting of 6-8 sporangia, medial or submarginal; indusia prominent, spherical, thinly membranous. n = 66.
On rocks in forests; 200-4000 m. Anhui, Guizhou, Hebei, Heilongjiang, Henan, Jiangxi, Jilin, Liaoning, Shandong, Sichuan, Zhejiang [Japan, Korea, Russia].