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FOC | Family List | FOC Vol. 2-3 | Davalliaceae | Davallia

5. Davallia solida (G. Forster) Swartz, J. Bot. (Schrader). 1800(2): 87. 1801.

阔叶骨碎补 kuo ye gu sui bu

Trichomanes solidum G. Forster, Fl. Ins. Austr. 86. 1786; Davallia solida var. latifolia Hooker; D. solida var. ornata Mettenius ex Kuhn; D. subsolida Ching; Humata solida (G. Forster) Desvaux; Stenolobus solidus (G. Forster) C. Presl.

Rhizome 4-14 mm in diam., densely covered with scales, generally not white waxy. Scales red-brown, with pale border from base to apex, narrowed evenly toward apex or above much broader base evenly narrowed toward apex, not or seldom curling backward, peltate, 5-10 × 1-1.2 mm, with multiseptate hairs at least when young (hairs at least at apex of young scales, ca. 1 mm, woolly). Stipe pale, adaxially grooved, 9-35 cm, glabrous or with few scales; lamina compound, bipinnate or tripinnate toward base and in middle part, deltoid and broadest toward base, 15-90 × 21-40 cm, glabrous (sometimes with hairs at junction of rachis and petiolule), not or slightly dimorphic. Longest petiolules 5-25 mm; pinnae linear-triangular or narrowly ovate, apex acuminate; longest pinnae 11-28 × 6-15 cm; pinnules of at least larger pinnae anadromous, deltoid or rhombic; longest pinnules 40-100 × 15-80 mm; ultimate pinnae linear-oblong or rhombic, lobed almost to midrib, or only shallowly lobed (in bipinnate fronds: ultimate segments shallowly lobed); ultimate segments 10-40 × 3-17 mm. Rachises and costae glabrous (often hairs at junction of petiolules). Margins of each pinna not thickened. Veins in sterile ultimate lobes pinnate, reaching margin or not; false veins not present. Sori separate, borne several on a segment, at forking point of veins; indusium also attached along sides, pouch-shaped, oblong, longer than wide, 1.2-2 × 0.5-1 mm, upper margin not elongated, truncate or slightly rounded, separated from or even with lamina margin; lamina not extending into teeth beyond a sorus.

Epiphytic, epilithic on different kinds of rocks, or terrestrial on different kinds of soils, in exposed places or in deep shade, from open rocky places and savannas to primary rain forests; sea level to 1500 m. Guangdong, Guangxi, Taiwan, Yunnan [Cambodia, India, Indonesia, Malaysia, Myanmar, Papua New Guinea, Philippines, Sri Lanka, Thailand, Vietnam (Tonkin); Pacific islands].

Sometimes the segments in Davallia solida are very narrow and the plant resembles D. solida var. fejeensis (Hooker) Nooteboom. Davallia solida is a widespread species and generally is easily recognizable by the black rhizome, which bears hairy scales that soon fall off leaving only the bases.


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