4. Humata griffithiana (Hooker) C. Christensen, Contr. U.S. Natl. Herb. 26: 293. 1931.
杯盖阴石蕨 bei gai yin shi jue
Davallia griffithiana Hooker, Sp. Fil. 1: 168. 1845; Davallia henryana Baker; D. platylepis Baker; D. tyermanii (T. Moore) Baker; Humata griffithiana var. tyermanii (T. Moore) Tagawa; H. henryana (Baker) Ching; H. platylepis (Baker) Ching; H. tyermanii T. Moore; Leucostegia griffithiana (Hooker) J. Smith.
Rhizome 3-6 mm in diam. (without scales), not white waxy. Scales whitish, brown, or red-brown, without pale border, narrowed evenly toward apex, curling backward or not, peltate, 6-9 × 1-1.5 mm, without multiseptate hairs, with marginal setae at least in distal part. Stipe adaxially grooved, 6-24 cm, glabrous or with few scales; lamina compound, bipinnate or tripinnate toward base and in middle part, deltoid and broadest toward base, 10-32 × 8-27 cm, glabrous, not or slightly dimorphic. Longest petiolules 2-10 mm; pinnae deltoid or linear; longest pinnae 6-16 × 4-8 cm; pinnules of at least larger pinnae anadromous, linear-oblong or narrowly ovate; longest pinnules 25-100 × 7-60 mm; ultimate pinnae narrowly ovate, lobed almost to midrib or only halfway toward midrib; ultimate segments or lobes obtuse or acute without a tooth, 2-5 × 2-3 mm. Rachises and costae glabrous. Margins of each pinna not thickened. Veins in sterile ultimate lobes simple, forked or pinnate, not reaching margin; false veins not present. Sori separate, borne several or single on a segment, at forking point of veins; indusium attached at base and only part of sides, semicircular, ± as wide as long, ca. 1 × 1-2 mm, upper margin not elongated, truncate or slightly rounded, separated from or even with lamina margin; lamina generally extending into a tooth at both sides or only at outside of a sorus, or not extending into teeth beyond a sorus.
Wet forests, climbing on tree trunks or rocks; near sea level to 2200 m. Fujian, Guangdong, Guangxi, Guizhou, Hunan, Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [Bhutan, India (Assam, Manipur), Japan (Okinawa), Laos, Myanmar (Kengtung), Vietnam (Tonkin)].
Humata henryana, H. platylepis, and H. tyermanii were treated as separate species in FRPS (6(1): 194-197. 1999), but comparisons of the types of Davallia henryana, D. platylepis, and H. tyermanii suggest that they all belong to the same species, H. griffithiana.