八角莲 ba jiao lian

Podophyllum versipelle Hance, J. Bot. 21: 362. 1883; P. esquirolii H. Léveillé.

Plants 40-150 cm tall. Rhizomes stout, with numerous fibrous roots; aerial stems erect, pale green, unbranched, glabrous. Leaves alternate; petiole of lower leaves 12-25 cm, petiole of upper leaves 1-3 cm; leaf blade suborbicular, to 30 cm in diam., thinly papery, abaxially pubescent, venation obviously raised, adaxially glabrous, palmately 4-9-lobed; lobes broadly triangular, ovate, or ovate-oblong, 2.5-4 cm, base 5-7 cm wide, margin remotely serrate, apex acute, not lobed. Inflorescence 5-8-fascicled flowers. Pedicel pendulous, slender, pubescent. Flowers attached near base of blade, red. Sepals oblong-elliptic, 0.6-1.8 cm × 3-8 mm, outside puberulent, inside glabrous, apex acute. Petals spatulate-obovate, ca. 2.5 × 0.8 cm, glabrous. Stamens ca. 1.8 cm; filaments shorter than anthers; anther connective slightly prolonged, glabrous, acute. Ovary ellipsoid, glabrous; style short; stigma shield-shaped. Berry ellipsoid or ovoid, ca. 4 × 3.5 cm. Seeds numerous. Fl. Mar-Jun, fr. May-Sep. 2n = 12*.

● Forests, thickets, shaded and wet places by streams, bamboo forests; 300-2400 m. Anhui, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangxi, Shanxi, Yunnan, Zhejiang.

Podophyllum versipelle was treated by Shaw (in Stearn, Gen. Epimedium, 276-282. 2002) as comprising two subspecies: subsp. versipelle ("Leaf lobes elobulate, upper leaf with 4-7 lobes on one side, lobes shallow 1/5-1/3 of radius; inflorescence 4-9 flowered; pedicels with hairs") and subsp. boreale J. M. H. Shaw ("Leaf lobes often with lobules, upper leaf with 4 or 5 lobes on one side, lobes deep, 1/2-2/3 of radius; inflorescence 4-10(-19) flowered; pedicels glabrous or rarely hairy").

Within Podophyllum versipelle subsp. boreale, Shaw (loc. cit.) recognized two varieties: P. versipelle var. boreale (lobes of leaves with convex margins, lobes entire or trilobulate; pedicels and leaves abaxially glabrous) and P. versipelle var. sichuanense J. M. H. Shaw (lobes of leaves with concave margins, lobes usually trilobulate; pedicels and leaves abaxially hairy).