58. Rubus sumatranus Miquel, Fl. Ned. India, Eerste Bijv. 307. 1860.
红腺悬钩子 hong xiao xuan gou zi
Rubus asper Wallich ex G. Don var. myriadenus (H. Léveillé & Vaniot) Focke; R. asper var. myriadenus (H. Léveillé & Vaniot) Focke subvar. grandifoliolatus (H. Léveillé) Focke; R. asper var. pekanius Focke; R. dolichocephalus Hayata; R. indotibetanus Koidzumi; R. myriadenus H. Léveillé & Vaniot; R. myriadenus var. grandifoliolatus H. Léveillé; R. rosifolius Smith subsp. sumatranus (Miquel) Focke; R. somae Hayata; R. sorbifolius Maximowicz; R. takasagoensis Koidzumi.
Shrubs erect or scandent. Branchlets brownish to dark reddish brown, cylindric, with long soft hairs, usually with scattered setose purplish red glandular hairs and curved prickles; glandular hairs and prickles unequal in length, glandular hairs to 4–5 mm, prickles to 8 mm. Leaves imparipinnate, to 15 cm, 5–7-foliolate, rarely 3-foliolate; petiole 3–5 cm, petiolule of terminal leaflet to 1 cm, lateral leaflets shortly petiolulate, petiolule and rachis soft hairy, with intermixed glandular hairs, with curved minute prickles; stipules lanceolate or linear-lanceolate, 6–8 mm, with soft hairs and intermixed glandular hairs; blade of leaflets ovate-lanceolate to lanceolate, 3–8 × 1.5–3 cm, abaxially villous, with gland-tipped hairs and small prickles along midvein, adaxially villous especially along midvein, base rounded, margin irregularly sharply serrate to doubly serrate, sometimes 3-lobed on terminal leaflet, apex acuminate. Inflorescences terminal, corymbose, 4–7 cm, 3- to several flowered, rarely flowers solitary; rachis and pedicels villous, with intermixed glandular hairs, with minute prickles; bracts lanceolate or linear, 5–7 mm, with soft hairs and intermixed glandular hairs. Pedicel 2–3 cm. Flowers 1–2 cm in diam. Calyx abaxially with soft hairs, with intermixed unequal long gland-tipped hairs; sepals reflexed in fruit, lanceolate or oblong-lanceolate, 7–12 × 2–4 mm; apex long caudate. Petals white, narrowly obovate or spatulate, slightly shorter than sepals, base clawed. Stamens many, shorter than petals. Pistils to 400; style and ovary glabrous; torus raised, oblong, base shortly stalked. Aggregate fruit orange-red, oblong, 1.2–1.8 cm × 7–11 mm, glabrous; pyrenes reticulate. Fl. Apr–Jun, fr. Jul–Aug. 2n = 14.
Forests, forest margins, thickets, bamboo forests, grasslands; 700--2500 m. Anhui, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [Bhutan, Cambodia, N India, Indonesia, Japan, Korea, Laos, Malaysia, Myanmar, Nepal, Sikkim, Thailand, Vietnam].