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FOC | Family List | FOC Vol. 13 | Violaceae | Viola

59. Viola betonicifolia Smith in Rees, Cycl. 37: Viola no. 7. 1817.

戟叶堇菜 ji ye jin cai

Viola betonicifolia subsp. dielsiana W. Becker; V. betonicifolia subsp. nepalensis (Gingins) W. Becker; V. caespitosa D. Don; V. inconspicua Blume subsp. dielsiana W. Becker; V. oblongosagittata Nakai var. violascens Nakai; V. patrinii Gingins var. caespitosa (D. Don) Ridley; V. patrinii var. laotiana H. Boissieu; V. patrinii var. nepaulensis Gingins.

Herbs perennial, acaulescent. Rhizome obliquely ascending or erect, usually short, 5-10 mm, stout, with several robust, long, yellowish roots. Leaves numerous, basal, rosulate; stipules dark brownish, lanceolate, 3-5 × ca. 1 mm, ca. 3/4 adnate to petiole, free part linear-lanceolate or subulate, margin entire or sparsely denticulate, apex acuminate; petiole long, 1.5-13 cm, narrowly and conspicuously winged in upper part, usually glabrous, sometimes puberulous in lower part; leaf blade narrowly lanceolate, narrowly triangular-hastate, or triangular-ovate, 2-7.5 × 0.5-3 cm, base truncate or shallowly cordate, sometimes broadly cuneate, apex acute, sometimes slightly obtuse; leaves accrescent after anthesis, both surfaces glabrous or subglabrous, lateral lobes spreading and conspicuously dentate, margin remotely shallowly crenate, more deeply dentate near base. Flowers white, light or deep purplish, deep-colored striate, 1.4-1.7 cm; pedicels long, equaling or exceeding leaves, slender, usually glabrous, sometimes puberulous in lower part, 2-bracteolate near middle; bracteoles linear. Sepals ovate-lanceolate or lanceolate, 5-6 × 1-1.6 mm, apex acuminate or ± acute, basal auricles shorter, 0.5-1 mm, 3-veined, margin narrowly membranous, apex rounded, sometimes remotely obtusely dentate. Upper petals obovate, 1-1.2 cm, bearded; lateral ones oblong-obovate, bearded, 1-1.2 cm; anterior one usually shorter, 1.3-1.5 cm (spur included); spur tubular, short, 1-2.5 mm, stout, 2-3 mm in diam., apex rounded, straight or slightly curved upward; anthers and appendage of connectives ca. 2 mm; spur of 2 anterior stamens 1-3 mm. Ovary ovoid-globose, ca. 2 mm, glabrous; styles clavate, slightly geniculate forward, gradually thickened in upper part; stigmas slightly thickened and narrowly margined on lateral sides and abaxially, shortly beaked in front, with a stigma hole at tip of beak. Capsule ellipsoid to oblong, 6-9 mm, glabrous. Seed ovoid, 1-1.2 mm. Fl. Feb-May, fr. May-Sep. 2n = 24*, 48, 72*.

Fields, roadsides, grasslands on mountain slopes, thickets, forest margins; below 1500(-2500) m. Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Taiwan, SE Xizang, Yunnan, Zhejiang [Afghanistan, Bhutan, Japan, India, Indonesia, Kashmir, Malaysia, Myanmar, Nepal, Philippines, Sri Lanka, Thailand, Vietnam; N Australia].


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