1 |
Plant generally wind-pollinated or self-pollinated, capitula small and not showy; all florets unisexual, or if bisexual then capitula aggregated into glomerules; ray florets absent |
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(2) |
+ |
Plant generally adapted for attracting pollinating insects, capitula colorful and attractive; some or all florets bisexual; ray florets present, sometimes inconspicuous, rarely absent |
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(4) |
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2 (1) |
Florets all bisexual; capitula aggregated into glomerules surrounded by leaflike bracts; achenes free and not surrounded by modified phyllaries into a bur or into spiny, tuberculate, or winged coat. |
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228 Lagascea |
+ |
Florets unisexual; capitula not aggregated into glomerules and lacking any outer leaflike bracts; achenes surrounded by modified phyllaries into a prickly bur or spines, tubercles, or wings |
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(3) |
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3 (2) |
Phyllaries in male capitula 1- or 2-seriate, free to base; phyllaries in female capitula 6-12-seriate, their distinct tips mostly ± hooked (distal 1-3 usually longer, stouter, and not hooked), whole becoming a hard, prickly perigynium or bur; florets 2. |
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232 Xanthium |
+ |
Phyllaries in male capitula 1-seriate, connate; phyllaries in female capitula 1-8-seriate, usually with free tips forming tubercles, spines, or wings; florets 1(-5). |
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233 Ambrosia |
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4 (1) |
Leaves and phyllaries with obvious oil glands |
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(5) |
+ |
Leaves and phyllaries lacking oil glands |
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(6) |
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5 (4) |
Phyllaries connate into a tube, sometimes splitting with age; style arms relatively long, spreading or arcuate; leaves lacking basal cilia. |
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202 Tagetes |
+ |
Phyllaries free; style arms minute, scarcely bifurcated; leaf bases distinctly ciliate. |
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203 Pectis |
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6 (4) |
Only ray florets fertile, ray achenes much longer than those of sterile disk florets |
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(7) |
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Disk florets fertile; ray florets present and fertile or sterile or absent |
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(9) |
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7 (6) |
Leaves alternate; ray achenes strongly flattened, each achene ± enclosed by and often attached to subtending phyllary and 2 adjacent paleae, all falling as a unit. |
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234 Parthenium |
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Leaves opposite; ray achenes thick, rounded or weakly compressed, without wings; achenes thick, not obviously flattened, not falling as unit with adjacent paleae |
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(8) |
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8 (7) |
Achenes enveloped and enclosed by prickly inner phyllaries. |
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214 Acanthospermum |
+ |
Achenes merely subtended by unarmed phyllaries. |
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217 Smallanthus |
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9 (6) |
Ray florets with or without short tubes, persistent with corollas fused to apex of achene. |
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211 Zinnia |
+ |
Ray florets deciduous, of if ray florets absent (i.e., capitula disciform or discoid) then corollas not fused to apex of achene |
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(10) |
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10 (9) |
Pappus of subulate to acerose scales, or spatulate, entire to erose, fimbriate, or laciniate, sometimes aristate, scales 1-seriate, or plumose, setiform scales (or flattened bristles) 1-seriate, or an apical peg |
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(11) |
+ |
Pappus absent, or awned |
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(13) |
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11 (10) |
Pappus of plumose setae. |
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212 Tridax |
+ |
Pappus of aristate or linear scales with erose or fimbriate margins, or absent |
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(12) |
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12 (11) |
Pappus of 6-12 aristate, rarely linear scales with erose margins; capitula > 10 mm in diam. |
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235 Gaillardia |
+ |
Pappus absent or of fimbriate, sometimes aristate scales; capitula 3-5 mm in diam. |
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213 Galinsoga |
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13 (10) |
Achenes compressed |
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(14) |
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Achenes all relatively plump, or 3-5-angled in ray florets and compressed in disk florets |
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(21) |
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14 (13) |
Pappus of retrorsely barbed awns; leaves opposite or upper alternate |
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(15) |
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Pappus absent, or persistent, of 2 bristly cusps or scales; leaves opposite |
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(17) |
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15 (14) |
Style branches with long hairs; pappus of 2 scabrid awns. |
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205 Glossocardia |
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Style branches with short minute papillae; pappus of 2-4 scabrid awns |
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(16) |
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16 (15) |
Anther filaments pubescent; achene apex beaked. |
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206 Cosmos |
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Anther filaments glabrous; achene apex narrow, not beaked. |
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207 Bidens |
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17 (14) |
Phyllaries 2-seriate, outer few and smaller, inner membranous, connate at base; pappus of 2-4 awns or scales or absent. |
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208 Coreopsis |
+ |
Phyllaries free, outer leathery, subequal, inner shorter, similar with paleae; ray achenes narrowly oval and winged, with 2 triangular scales |
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(18) |
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18 (17) |
Achenes markedly dimorphic, those of ray florets with conspicuous, lacerate winged margin, others wingless. |
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219 Synedrella |
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Achenes all ± alike, wingless or somewhat thickly winged distally, not lacerate |
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(19) |
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19 (18) |
Shrubs or small trees; synflorescence of numerous capitula in congested panicles or corymbs; achenes wingless. |
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223 Clibadium |
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Annual or perennial herbs; synflorescence of solitary capitula or capitula several in dense clusters; achenes winged or angled |
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(20) |
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20 (19) |
Capitula radiate; pappus of 2 stout awns. |
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220 Calyptocarpus |
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Capitula discoid; pappus peglike. |
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221 Eleutheranthera |
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21 (13) |
Achenes enclosed by inner phyllaries or outer paleae |
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(22) |
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Achenes not enclosed by inner phyllaries |
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(24) |
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22 (21) |
Capitula tightly or loosely aggregated on short peduncles in corymbose panicles. |
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204 Flaveria |
+ |
Capitula axillary or in sparse cymes or panicles |
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(23) |
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23 (22) |
Female florets 1-seriate; phyllaries 5, spreading; capitula in sparse panicles, peduncles long. |
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216 Sigesbeckia |
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Female florets multiseriate; phyllaries 4, broad, in 2 pairs; capitula axillary, peduncles very short. |
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209 Enydra |
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24 (21) |
Paleae narrow, long, flat; ray florets 2-seriate, lamina small; pappus absent or of 2 short awns |
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(25) |
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Paleae concave or folded, ± enclosing florets |
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(26) |
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25 (24) |
Ray floret lamina white; achene body tuberculate. |
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222 Eclipta |
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Ray floret lamina yellow; achene body striate. |
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215 Guizotia |
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26 (24) |
Achenes in ray florets broadly ovate or elliptic, 3-angled; disk achenes ellipsoid, strongly compressed; pappus absent or of up to 10 awnlike bristles. |
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210 Acmella |
+ |
Achenes in bisexual florets 4- or 5-angled, or compressed |
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(27) |
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27 (26) |
Ray florets sterile |
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(28) |
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Ray florets fertile |
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(31) |
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28 (27) |
Paleae completely investing and falling with accompanying achene, each forming a hardened perigynium. |
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227 Sclerocarpus |
+ |
Paleae sometimes conduplicate, ± enfolding achenes, not forming perigynium |
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(29) |
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29 (28) |
Pappus absent, a small crown of minute scales, or of 2-4 small scales; receptacles conical to columnar. |
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229 Rudbeckia |
+ |
Pappus of 2 awns sometimes with a few scales, or a crown of fused scales, or of 2 awns with fused scales, or absent; receptacles flat to shallowly convex |
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(30) |
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30 (29) |
Peduncles usually distally dilated, always fistulose; pappus absent or coroniform, of connate scales, 1 or 2 scales sometimes subulate to aristate. |
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230 Tithonia |
+ |
Peduncles never fistulose; pappus absent or readily deciduous, of 2(or 3) usually lanceolate, aristate, or erose scales, plus 0-8 usually shorter scales. |
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231 Helianthus |
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31 (27) |
Pappus elements 2-5, unequal, spinelike or squamalike, persistent, base connate; ray florets female, ray floret lamina short or very short, apex 2-4-dentate; capitula small. |
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218 Blainvillea |
+ |
Pappus absent, or scalelike, cyathiform, coroniform, or of 1 or 2 setae; capitula relatively large |
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(32) |
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32 (31) |
Corollas orange to yellow; outer phyllaries herbaceous and larger than inner; leaves sessile or very shortly petiolate; plants mainly prostrate; achenes triangular (ray) or compressed (disk); capitula always solitary, terminal (but appearing axillary) on erect peduncles; achene beak and pappus obscured at maturity by corky collar. |
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224 Sphagneticola |
+ |
Corollas white or yellow; outer phyllaries ± equal in size to inner; leaves usually conspicuously petiolate; plants mostly erect or ascending [sometimes vines or subshrubs outside of our area]; synflorescence of 1-3(-6) capitula, terminal or axillary; pappus of obvious setae (rarely absent) |
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(33) |
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33 (32) |
Ray florets female and sterile or neuter; pappus awns caducous. |
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225 Melanthera |
+ |
Ray florets female and fertile; pappus absent or single awn persistent. |
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226 Wollastonia |
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