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FOC | Family List | FOC Vol. 2-3 | Dryopteridaceae | Dryopteris

79. Dryopteris carthusiana (Villars) H. P. Fuchs, Bull. Soc. Bot. France. 105: 339. 1959.

刺叶鳞毛蕨 ci ye lin mao jue

Polypodium carthusianum Villars, Hist. Pl. Dauphiné 1: 292. 1786.

Plants 30-60 cm tall. Rhizome erect or obliquely ascending, short, stout, scaly; scales pale brown, ovate or ovate-lanceolate, membranous, dentate. Fronds caespitose; stipe stramineous, as long as lamina, longitudinally grooved, with ovate, dentate scales; lamina deltoid or ovate-oblong, 20-30 cm, 15-16 cm wide at lower part, bipinnate-pinnatifid; pinnae 12 or 13 pairs, oblique, basal pair largest, shortly stalked, upper ones gradually shortened, connected by wing at base, lowest ones deltoid, 9-10 × 4-5 cm, base subtruncate, shortly stalked, apex acuminate; pinnules 8 or 9 pairs, basal pair largest, upper ones gradually shortened, connected by broad wing, basal pair opposite or subopposite, deltoid, 2.5-3 cm, base 1-1.5 cm wide, shortly stalked; acroscopic pinnules smaller than basiscopic ones, nearly at right angle to costae, basiscopic one parallel to pinna rachis or slightly oblique, pinnatifid; segments 4 or 5 pairs, basal pair asymmetrical, basiscopic one smaller than others, oblique, acroscopic one oblong, 8-10 × ca. 2 mm, almost parallel to costa, aristate. Veins impressed adaxially, raised abaxially, pinnate, simple, extending into each tooth. Sori terminal on apex of lateral veins, in 2 rows along main vein of pinnules; indusia pale brown, membranous, with an incised margin, often deciduous at maturity.

Montane forests, open forests; ca. 2000 m. Xinjiang (Zhaosu) [Europe, E North America].

According to the treatment by Chang Y. Yang (Fl. Xinjiang. 1: 46. 1992), Dryopteris carthusiana was collected from Zhaosu, Xinjiang, in 1974. This collection has not been seen by the present authors.


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