钩距黄堇 gou ju huang jin

Corydalis binderae Fedde subsp. pseudohamata (Fedde) Z. Y. Su; C. fluminicola W. W. Smith; ?C. hamata var. ramosa Z. Y. Su; C. pseudohamata Fedde.

Herbs, perennial, 15-40(-60) cm tall, glabrous. Rootstock 2-3 cm, stout, with long soft fleshy roots 1-2 mm in diam., with 2-sided apical leaf rosette. Stems radiating, 3-12, decumbent at base, ascending (in shaded subalpine places erect), branched, with 4-7(-10) leaves in upper 2/3. Rosette leaves several; petiole 5-11 cm, long vaginate with membranous margin; blade ob-long, 7-15(-20) × 3-6(-10) cm, bipinnate; pinnae in 3-5 pairs, nearly sessile or lower sometimes very long stalked; pinnules usually 3, deeply 3-6-lobed; segments lanceolate to oblong, 3-6 × 2-3 mm. Cauline leaves like basal leaves, lower petiolate, upper sessile; blade 3-10 × 2-4 cm, with 2-4 pairs of leaflets. Raceme subspicate, very dense, 20-40-flowered, 4-10 cm; flowers often nutant; bracts obovate to lanceolate, 5-10(-15) mm, entire or lower (rarely most) ones often divided, sometimes with purplish margin. Pedicel erect, 5-10 mm. Sepals early withering, turning brown, broadly ovate, 2-3(-4) × 2-3 mm, finely and shallowly dentate (NE) to deeply laciniate dentate (SW). Corolla yellow or cream with yellow tip, sometimes with a purplish suffusion on spur, rarely purple with white apex; inner petals with blue dorsal crests, rarely dark purple. Outer petals rounded, obtuse to subacute, with 1-2-mm-wide entire crest usually overtopping apex; upper petal 16-20 mm; spur strongly curved in a tight semicircle, apex thickened, 4-6 mm (measured in middle of spur); nectary ca. 1/2 as long as spur; lower petal with narrow claw; inner petals 11-13 mm, with swollen dorsal crests overtopping apex. Stigma square, emarginate, with 4-6 marginal simple papillae; geminate papillae lateral (sometimes missing) and in basal corners (sometimes missing). Capsule lanceolate, ca. 15 × 4-5 mm, 8-14-seeded; style 3.5-4 mm. Seeds in 2 rows, 1.8-2.1 mm, almost smooth (finely reticulate under strong lens). Fl. and fr. Jul-Sep.

● Wet alpine scree, brook sides in alpine meadows, often in running water; 3300-5200 m. W Sichuan, E Xizang (Jomda, Sogxian, Zogang), NW Yunnan.

Plants at lower elevations in shade tend to be coarse and erect with strict erect branches. Plants in the NE part of the distribution area (NE from Yajiang, W Sichuan) tend to have much less deeply dentate sepals. A form from Daocheng (SW Sichuan) and Zogang (E Xizang), from open alpine rocky brooks, has deeply divided bracts and dark purple flowers white at apex.