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FOC | Family List | FOC Vol. 7 | Papaveraceae | Corydalis

202. Corydalis linarioides Maximowicz, Bull. Acad. Imp. Sci. Saint-Pétersbourg, sér. 3. 24: 27. 1878.

条裂黄堇 tiao lie huang jin

?Corydalis linarioides var. fissibracteata Fedde; ?C. schlechteriana Fedde.

Herbs, perennial, 10-50 cm tall, glabrous or sparsely papillose at base of stem and sometimes abaxially on leaf; stems and petioles of radical leaves attenuate to filiform underground base. Storage roots few to several, spindle-shaped, 1-2 cm, narrowed into short or indistinct stalk (more distinct on young tubers). Stems 1-5, usually simple, rarely with 1 or 2 branches, with 2 or 3 leaves in upper 1/2. Radical leaves few; petiole to 14 cm; blade 2-4 × 2-5 cm, pinnate; leaflets cut to base into 2 or 3 linear to oblong lobes. Cauline leaves sessile; blade pinnate; pinnae 2-4 pairs, linear, 2-5 cm × 1-5 mm, entire. Raceme 6-20-flowered, dense, elongating in fruit; bracts lanceolate, entire or lower and middle bracts deeply pectinate-dentate to palmatifid, equaling pedicels. Pedicel 5-10 mm in flower, 10-15 mm and reflexed in fruit. Sepals whitish, 0.3-1 × 0.3-2 mm, finely and sharply dentate. Petals yellow, usually with distinct darker veins, apex usually orange, dorsal crest of inner petals often with small brown mark; upper petal 16-23 mm, subacute to subobtuse, abaxial crest 1-2 mm wide, from apex extended to spur, sometimes not reaching apex; spur straight or slightly downcurved at apex, cylindric, 8-15 mm; nectary ca. 1/2 as long as spur; lower petal obovate, 9-11 mm, abaxial crest smaller than that of upper petal; inner petals 7-9 mm. Stigma square; simple marginal papillae: median 2 distinct, lateral broad and indistinct; geminate papillae lateral (submarginal) and terminating well-developed basal lobes. Capsule narrowly oblong, 10-14 × 1.5-2 mm, smooth or sometimes with thin papillose lines, 3-8-seeded. Seeds 1-ranked, ca. 1.5 mm, smooth, with rather large elaiosomes. Fl. and fr. Jun-Sep.

● ?Forests, forest margins, shrubs, meadows, alpine pastures; (2100-)3200-4500 m. Gansu, Ningxia (Guyuan), Qinghai, Shaanxi, Shanxi, Sichuan, Xizang.

As here circumscribed, this taxon is probably heterogeneous. At least the following two forms can be recognized, but more research is needed to establish their status. The capsule and seed characters in the above description refer to Type B.

Type A. Stem robust, 10-25 cm, simple, glabrous; leaves stiff, adaxially dark glossy green, abaxially frequently with coarse 1-celled hairs on veins, leaflets 1-2 mm wide; lower and middle bracts pectinate-dentate with long narrow teeth; spur 10-15 mm. Open areas. This corresponds to Corydalis linarioides var. fissibracteata Fedde, p.p., and is similar to C. eugeniae in general appearance.

Type B. Stem slender, 20-50 cm, simple or rarely branched, glabrous or sometimes papillose-hairy at base; leaves soft, slightly glaucous, glabrous, leaflets 1-5 mm wide; bracts lanceolate, entire; spur 8-11 mm. More shaded places, e.g., in shrubs. This possibly corresponds to the type of Corydalis linarioides.

The complex comprising Corydalis linarioides, C. eugeniae, C. atuntsuensis, and C. mayae is far from satisfactorily elucidated, and the variation is reticulate. We have here taken storage root form as a primary discriminant: a) oblong with distinct thin long branched stalks (C. eugeniae); b) short fusiform with indistinct or short thick stalks (C. linarioides); c) long narrowly fusiform, gradually attenuate to base (C. atuntsuensis); d) similar, but stalks narrower (C. mayae).


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