东方荚果蕨 dong fang jia guo jue

Struthiopteris orientalis Hooker, Sec. Cent. Ferns, t. 4. 1861; Matteuccia japonica (Hayata) C. Christensen; M. orientalis (Hooker) Trevisan; Onoclea orientalis (Hooker) Hooker; Pentarhizidium japonicum Hayata; Pteretis japonica (Hayata) Ching; P. orientalis (Hooker) Ching; Pterinodes orientalis (Hooker) Kuntze.

Rhizome shortly creeping, apex ascending, dark brown, apical part densely covered with brown scales; scales lanceolate, ca. 2 cm, membranous, entire, apex fibriform. Fronds approximate. Stipe of sterile frond brown, 30-70 × 0.3-1 cm, base swollen, triangular, bearing short pneumatophores, densely scaly, upward sparsely scaly; sterile lamina bipinnatifid, ovate-deltoid, narrowed upward, slightly narrowed downward, 40-82 × 18-40 cm, thinly papery, with sparse fibriform scales on rachis and costae; pinnae 15-20 pairs, alternate, oblique, 2.5-3 cm apart, basal pinnae as long as above, base slightly narrowed, sessile, apex acuminate; lobes narrowly hemielliptic, oblique, entire or slightly dentate, apex obtuse or acute; veins obvious, pinnate, veinlets simple or occasionally furcate, reaching lamina margin. Stipe of fertile frond 20-45 cm; fertile lamina as long as or shorter than sterile lamina, pinnate, elliptic, 12-40 × 5-10 cm; pinnae oblique, close, dark purple, linear, ca. 10 × 0.5 cm, hardened. Sori fused into linear coenosori when mature; indusium membranous, fixed at posterior end, free distally. 2n = 80, 82.

Streams in dense forests, slopes at forest margins, semi-shaded places in forests; 1000-2700 m. Anhui, Chongqing, Gansu, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangxi, Shanxi, Sichuan, Taiwan, Xizang, Zhejiang [India, Japan, Korea, Russia].

Matteuccia orientalis f. monstra Ching & K. H. Shing (J. Arnold Arbor. 64(1): 25. 1983) was described from Hubei.