疣状叉蕨 you zhuang cha jue

Phlebiogonium impressum Fée, Mém. Foug. 5: 314. 1852; Aspidium immersum Hooker (1862), not Blume (1828); A. prominens Alderwerelt; A. variolosum Wallich ex Hooker; A. zollingerianum Beddome (1868), not Kunze (1846); Nephrodium variolosum (Wallich ex Hooker) Baker; Sagenia variolata T. Moore; Tectaria prominens (Alderwerelt) C. Christensen; T. variolosa (Wallich ex Hooker) C. Christensen.

Plants terrestrial, 40-80 cm tall. Rhizome creeping to suberect, short, stout, densely scaly at apex and stipe bases; scales stiff, brown, linear-lanceolate, 6-8 mm, membranous, ciliate, apices long acuminate. Fronds clustered, subdimorphic, fertile fronds rather tall but narrowed; stipe stramineous, 30-50 cm, throughout covered with light brown hairs. Sterile lamina bipinnatifid to tripinnatifid, deep green or brown when dried, pentagonal, 30-35 × 30-35 cm, thickly herbaceous to leathery, both surfaces pubescent; rachises and costae stramineous, densely covered with articulate hairs; terminal pinna pinnatifid, base cuneate to decurrent, apex acuminate; lateral pinnae 1-5 pairs, opposite, interval 4-6 cm, oblique; basal pinnae pinnate with 1-3 undulate or pinnatifid lobes, 15-20 cm, rather large, stalks 2-3 cm, lower lobes larger than upper ones; middle pinnae falcate-lanceolate, undulate to pinnatifid, stalks becoming shorter, oblique, 6-8 cm, apices long caudate. Veinlets forming copious subhexagonal areoles with cross veins, included veinlets simple or not, distinctly forming long areoles with arcuate veinlet beside pinna rachis and main veins. Sori orbicular, terminal on veinlets, in one 1 row beside main veins; indusia brown, orbicular or reniform, firm, persistent.

Usually on limestone in dense forests; 100-500 m. Guangdong, Guangxi, Guizhou, Hainan, Taiwan, Yunnan [India, Indonesia, Laos, Malaysia, Nepal, Thailand, Vietnam].

Knapp (Ferns Fern Allies Taiwan, 961. 2011) included Aspidium subtriphyllum (Hooker & Arnott) Hooker f. cuspidatopinnatum Hayata (Icon. Pl. Formosan. 4: 189. 1914; Aspidium cuspidatopinnatum (Hayata) Ching; Tectaria cuspidatopinnata (Hayata) C. Christensen) within T. impressa.