8. Damnacanthus macrophyllus Siebold ex Miquel, Ann. Mus. Bot. Lugduno-Batavi. 3: 110. 1867.
浙皖虎刺 zhe wan hu ci
Damnacanthus indicus C. F. Gaertner f. macrophyllus (Siebold ex Miquel) Makino; D. indicus var. macrophyllus (Siebold ex Miquel) Makino; D. indicus var. parvispinus Koidzumi; D. major Siebold & Zuccarini var. macrophyllus (Siebold ex Miquel) Maximowicz; D. major var. parvispinus (Koidzumi) Koidzumi; D. major var. submitis Maximowicz ex Regel; D. moniliformis Koidzumi; D. minutispinus Koidzumi; D. shanii K. Yao & M. B. Deng.
Shrubs, 1-2 m tall. Roots fleshy, moniliform. Branches puberulent to hispidulous, with 8 alternately thick and thin striae, with few deciduous or persistent spines 2-6 mm. Petiole of developed leaves 1-2 mm, glabrous or sparsely puberulent to puberulent or hirtellous; leaf blade drying stiffly papery, ovate to oblong-ovate, lanceolate, or oblong-lanceolate, 3-6(-8) × 1-2.5(-3) cm, adaxially glabrous, abaxially glabrous or puberulent along veins, base cuneate to rounded, margins flat to thinly revolute, apex shortly acuminate or acute; midrib thinly prominent adaxially; secondary veins 3 or 4(-7) pairs; stipules persistent, interpetiolar, triangular, 0.3-1 mm, puberulent to glabrescent, acute to glandular-multifid or -fimbriate. Inflorescences strigillose to puberulent. Pedicels 1-2 mm. Calyx strigillose to glabrescent; hypanthium portion obconic, 1.2-1.5 mm; limb 1-1.5 mm, lobed for 1/3-1/2; lobes triangular to ovate. Corolla white, outside glabrous; tube 8(-13) mm; lobes ovate-triangular, ca. 2 mm. Drupes ca. 5 mm in diam., with pedicels sometimes elongated to 5 mm. Fl. Apr-Jun, fr. Oct-Dec.
Streamsides in sparse or dense forests on mountains; 800-1000 m. Anhui, Fujian, Guangdong, Guizhou, Yunnan, Zhejiang [Japan].
Naiki and Nagamasu (Amer. J. Bot. 91: 664-671. 2004) reported the breeding biology of this species as monomorphic with "pin" type flowers.
"Damnacanthus subspinosus var. salicifolius" (M. B. Deng & K. Yao, Bull. Bot. Res., Harbin 10(4): 2. 1990) belongs here but was not validly published because two gatherings were designated as types (Vienna Code, Art. 37.2).