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7f. Galium boreale var. kamtschaticum (Maximowicz) Nakai, Bot. Mag. (Tokyo). 23: 103. 1909.

堪察加拉拉藤 kan cha jia la la teng

Galium boreale f. kamtschaticum Maximowicz, Mém. Acad. Imp. Sci. St.-Pétersbourg Divers Savans 9 [Prim. Fl. Amur.]: 141. 1859; G. boreale var. koreanum Nakai.

Leaf blade broadly lanceolate or ovate-lanceolate, 6-10 mm wide, abaxially sparsely hairy or scabrous at least along veins. Ovary and mericarps densely hirsute or tomentose. Fl. and fr. Jun-Sep.

Mountain slopes, farmland sides, riversides, river valleys, grasslands, meadows; 800-2400 m. Heilongjiang, Henan, Jilin, Liaoning, Nei Mongol, Shaanxi, Shanxi, Sichuan, Xinjiang [Kashmir, Korea, Mongolia, Russia; C Asia ("Turkestan"), NE Europe].

Nakai (Bot. Mag. (Tokyo) 23: 103. 1909) commented on Japanese plants and presented a key to Galium, including "Galium boreale var. kamtschaticum Maximowicz" with no further information. Some authors have considered this a validly published combination (e.g., in Govaerts et al., World Checkl. Rubiaceae;; accessed on 16 Nov 2007), whereas others have not (e.g., W. C. Chen, loc. cit.: 263) and have instead cited J. Coll. Sci. Imp. Univ. Tokyo 31: 498. 1911 (sometimes called Fl. Koreana 2) as the place of publication. Nakai’s new combination was indeed validly published in 1909 because, before 1953, merely citing an author’s name could constitute indication of a basionym (Vienna Code, Art. 33.2). Pobedimova et al. (loc. cit.: 419) erroneously attributed this varietal name to Maximowicz, 1859; it was published as a "forma" there.


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