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FOC | Family List | FOC Vol. 11 | Celastraceae | Euonymus

2. Euonymus sect. Echinococcus Nakai, J. Jap. Bot. 17: 617. 1941.

刺果卫矛组 ci guo wei mao zu

Shrubs or small trees, deciduous or evergreen. Flowers 4-merous, rarely 5-merous. Capsule with tubercles or prickles, 4- or 5-lobed at maturity.

Eighteen species: E, S, and SE Asia; 14 species (ten endemic) in China.


1 Capsule with tubercles (not prickles), entire surface densely tuberculate.   28 E. verrucocarpus
+ Capsule with prickles, not tubercles   (2)
       
2 (1) Capsule with needle to threadlike prickles, sparsely covering surface   (3)
+ Capsule with sturdy prickles, densely covering surface   (7)
       
3 (2) Prickles fine and slender or sometimes needlelike; capsule large, 1.3-1.8 cm in diam..   25 E. lushanensis
+ Prickles straight, not fine and slender; capsule small, less than 1 cm in diam   (4)
       
4 (3) Leaf blade entire.   26 E. potingensis
+ Leaf blade crenulate to serrate   (5)
       
5 (4) Dwarf shrubs, under 30 cm tall.   19 E. aculeolus
+ Shrubs, more than 1 m tall   (6)
       
6 (5) Leaf blade 5-12 × 3-4.5 cm, veins impressed adaxially, prominent abaxially; capsule ca. 8 mm in diam..   23 E. chui
+ Leaf blade 4-6 × 2-3 cm, veins invisible; capsule 8-10 mm in diam..   27 E. spraguei
       
7 (2) Capsule grayish green or gray when dry, prickles more than 1 cm, flattened at base.   17 E. actinocarpus
+ Capsule brown, yellow, tawny, black, or purple when dry, prickles less than 1 cm   (8)
       
8 (7) Inflorescences more than 10 cm; flowers more than 10   (9)
+ Inflorescences less than 9 cm; flowers less than 9   (10)
       
9 (8) Leaf blade ovate or ovate-elliptic, adaxially wrinkled, petiole 5-8 mm.   20 E. balansae
+ Leaf blade oblong to elliptic, adaxially never wrinkled, petiole 1-2 cm.   16 E. acanthocarpus
       
10 (8) Leaf blade pubescent on both sides.   21 E. acanthoxanthus
+ Leaf blade glabrous   (11)
       
11 (10) Leaves sessile or petiole less than 4 mm   (12)
+ Leaf petiole more than 8 mm   (13)
       
12 (11) Leaf blade less than 4 × 2 cm.   22 E. chenmoui
+ Leaf blade ca. 5 × 2-3 cm.   24 E. echinatus
       
13 (11) Leaf blade leathery, 7-10 × 3-6 cm; capsule 1-1.5 cm in diam..   18 E. aculeatus
+ Leaf blade thinly leathery or papery, 10-15 × 2.5-4.5 cm; capsule 1.5-2 cm in diam..   29 E. wilsonii

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