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FOC | Family List | FOC Vol. 20-21 | Asteraceae | Artemisia

2. Artemisia sect. Abrotanum (Duhamel du Monceau) Besser, Bull. Soc. Imp. Naturalistes Moscou. 1: 222. 1829.

艾蒿组 ai hao zu

Abrotanum Duhamel du Monceau, Traité Arbres Arbust. 1: 19. 1755.

Herbs, annual or perennial, rarely subshrubs, pubescent, tomentose, or glabrous, rarely arachnoid, without glandular or viscid hairs (sessile glands often present). Leaves 2-4-pinnatisect; lobules pectinate, filiform, narrowly linear, or lanceolate, less than 1(-1.5) mm wide. Capitula hemispheric, globose, or subglobose, rarely ovoid. Phyllary margins scarious, midvein green or sometimes colored. Receptacle glabrous. Marginal florets (1-)3-20; corolla narrowly tubular or rarely narrowly conical, 2- or 3(or 4)-toothed or without teeth. Disk florets (7-)10-80, bisexual, ovaries well developed; style ca. as long as or longer than corolla, branches divergent, recurved at apex.

About 65 species: N Hemisphere, mostly in Asia and Europe, a few species extending into N Africa, Oceania, and the Americas; 30 species (seven endemic) in China.

Most species contain mainly monoterpenoids and also a few sesquiterpenoids. This section has been confused with Artemisia sect. Artemisia.


1 Leaves pinnatisect, with ultimate lobules filiform, linear-filiform, filiform-clavate, narrowly lanceolate, or linear-lanceolate, (5-)10+ × 0.5-1(-1.5) mm, apex acute   (2)
+ Leaves pinnatipartite or pinnatisect, with ultimate lobules pectinate, serrate, or smoothly obtusely lacerate, less than 5(-8) × 1.5(-2) mm, or lobules minute but then lobes 1-2 mm wide with obtuse apex   (9)
       
2 (1) Herbs annual, biennial, or short-lived perennial; rootstock a simple taproot, sometimes ± fusiform   (3)
+ Herbs perennial, subshrubs, or shrubs; rootstock often branched, ± woody   (6)
       
3 (2) Herbs biennial or short-lived perennial; capitula obconical, 4-5 mm in diam.   52 A. fukudo
+ Herbs annual; capitula ovoid or subglobose, 2-3 mm in diam   (4)
       
4 (3) Middle stem leaves pinnatisect, segments 2(or 3) pairs; plants pubescent, ± glabrescent.   53 A. przewalskii
+ Middle stem leaves 2- or 3-pinnatisect, segments (2 or)3 or 4 pairs; plants glabrous or nearly so   (5)
       
5 (4) Stem branches short or absent; middle stem leaves (1 or)2-pinnatisect, segments (2 or)3 or 4 pairs; capitula 2-10 grouped, in narrow panicles.   46 A. palustris
+ Stem branches divaricate; middle stem leaves 2- or 3-pinnatisect, segments 4 or 5 pairs; capitula 2-5 grouped, in broad lax panicles.   47 A. aurata
       
6 (2) Middle stem leaves 2- or 3-pinnatisect, lobules narrowly linear, densely silvery gray or yellowish sericeous-tomentose; corolla limb puberulent.   49 A. austriaca
+ Middle stem leaves 1- or 2-pinnatisect, lobules linear, filiform, or filiform-clavate, pubescent or arachnoid pubescent, sometimes glabrescent later, or densely white tomentose abaxially; corolla limb glabrous   (7)
       
7 (6) Stem much branched; middle stem leaves 2-4 cm, 2(or 3)-pinnatisect; abaxial leaf surface and phyllaries densely white tomentose.   50 A. brachyloba
+ Stem shortly capitellate branched or not; middle stem leaves 0.8-2.5(-3) cm, 1- or 2-pinnatisect; leaves and phyllaries puberulent or leaves glabrescent   (8)
       
8 (7) Stem terete; lobules of middle stem leaves filiform, apex acute.   44 A. adamsii
+ Stem 4-ribbed; lobules of middle stem leaves filiform-clavate or linear-filiform, apex obtuse.   45 A. dalai-lamae
       
9 (1) Herbs annual or biennial; stem solitary, more than 20 cm; rootstock a simple taproot, sometimes ± fusiform   (10)
+ Herbs perennial or subshrubs; stems several or many, rarely solitary; roots usually woody, or herbs dwarf, less than 20 cm and with thick stock and ± fusiform root   (15)
       
10 (9) Leaves ± succulent, middle stem leaves 1- or 2-pinnatisect, segments 1 or 2 pairs, with obtuse apex, entire or 1-3-crenate or cleft.   51 A. succulenta
+ Leaves thinly papery, middle stem leaves 2- or 3-pinnatisect, segments 2 or more pairs, lobes or lobules acute or obtuse apically   (11)
       
11 (10) Basal leaves many, forming rosettes, segments more than 20 pairs; phyllaries with brown or purple-brown scarious margins; marginal female florets 3-8.   41 A. hedinii
+ Basal leaves not forming distinct rosettes, segments 3-8(-10) pairs; marginal female florets 10-12   (12)
       
12 (11) Involucre 5-7 mm in diam.; phyllaries puberulent; disk florets 80-150; middle stem leaf segments 3 or 4 pairs, lobes serrate.   38 A. shangnanensis
+ Involucre 1.5-5(-7) mm in diam.; phyllaries glabrescent; disk florets 10-40; middle stem leaf segments 4-10 pairs, lobes pectinate   (13)
       
13 (12) Lower stem leaf petioles 2-6 cm; leaf blades minutely and lanceolately pectinate; synflorescence narrow and spikelike.   42 A. tournefortiana
+ Lower stem leaf petioles less than 2 cm; leaf blades pectinate; synflorescence broader, often a conical, compound panicle   (14)
       
14 (13) Middle stem leaves abaxially gray-yellowish or yellowish and gland-dotted; leaf rachis usually not pectinate, midvein obvious, slightly prominent; capitula 1.5-2.5 mm in diam.   40 A. annua
+ Middle stem leaves abaxially green, not gland-dotted; leaf rachis pectinate, midvein flat; capitula 3.5-5(-7) mm in diam.   39 A. caruifolia
       
15 (9) Herbs dwarf perennial, main root ± fusiform; stem usually less than 15 cm tall; leaves 1(or 2)-palmatisect, segments 1 or 2 pairs, without lobules or occasionally with 2 or 3 minute lobules; synflorescence spikelike.   48 A. caespitosa
+ Herbs perennial or subshrubs, roots woody; stem more than 15 cm tall; leaves 1(or 2)-pinnatisect, segments 3 or more pairs; synflorescence paniculate, racemelike, or spikelike   (16)
       
16 (15) Capitula broadly ovoid; phyllaries gray arachnoid pubescent; lobes of leaves suborbicularly and obtusely serrate.   31 A. baimaensis
+ Capitula hemispheric, globose, or subglobose; phyllaries puberulent, later glabrescent; lobes of leaves pectinate, or lobules lanceolate or linear-lanceolate, rarely linear-falcate   (17)
       
17 (16) Middle stem leaves 1- or 2-pinnatipartite, lobules pectinate   (18)
+ Middle stem leaves 2- or 3(or 4)-pinnatisect, lobules serrate   (19)
       
18 (17) Subshrubs; stem more than 70 cm; lobules of leaves elliptic or lanceolate and serrate; capitula 4-6 mm in diam.   32 A. medioxima
+ Herbs perennial; stem 20-70 cm; lobules of leaves pectinate; capitula 3-4 mm in diam.   35 A. latifolia
       
19 (17) Capitula hemispheric, rarely subglobose, 4-7 mm in diam., in racemes, spikes, or panicles   (20)
+ Capitula subglobose, 2.5-4 mm in diam., in panicles   (25)
       
20 (19) Herbs perennial; corolla limb puberulent   (21)
+ Subshrubs; corolla limb often glabrous   (22)
       
21 (20) Middle stem leaves pinnatisect, segments 5-7(or 8) pairs, lobes pectinate or deeply serrate; phyllaries with entire broad brown scarious margins.   36 A. phaeolepis
+ Middle stem leaves 2-pinnatisect or -partite, segments 4(or 5) pairs, lateral mid-lobes deeply 2-4-serrate; phyllaries with lacerate brown scarious margins.   43 A. tsugitakaensis
       
22 (20) Petioles of middle stem leaves 6-16 cm; leaves 1(or 2)-pinnatisect, lobes minutely irregularly pectinate-serrate.   34 A. maximovicziana
+ Petioles of middle stem leaves less than 3 cm; leaves 2 or 3(or 4)-pinnatisect, lobules minutely serrate   (23)
       
23 (22) Middle stem leaves ± sessile, 2(or 3)-pinnatisect, lobules minutely linear or linear-lanceolate or falcate, 0.5(-1) mm wide; capitula sessile or shortly pedunculate; phyllaries pubescent.   24 A. macrantha
+ Middle stem leaves petiolate, 2 or 3(or 4)-pinnatisect, lobules minute, pectinate, linear, elliptic, or ovate, (0.5-)1-1.5 mm wide; capitula pedunculate; phyllaries glabrescent or puberulent   (24)
       
24 (23) Middle stem leaves 3(or 4)-pinnatisect, segments 5-8 pairs, abaxial midvein obviously convex; peduncles 2-4 mm; phyllaries glabrescent.   26 A. polybotryoidea
+ Middle stem leaves 2(or 3)-pinnatisect, segments 3(or 4) pairs, abaxial midvein not convex; peduncles 4-15 mm; phyllaries puberulent.   27 A. aksaiensis
       
25 (19) Middle stem leaves with 6-8 pairs of segments, abaxially white tomentulose.   37 A. tanacetifolia
+ Middle stem leaves with 3-5 pairs of segments, abaxially pubescent or glabrescent   (26)
       
26 (25) Plants with short, woody vegetative shoots and slender, virgate synflorescences; petiole of middle stem leaves 0.3-0.5 cm; capitula with 16-20 disk florets.   33 A. freyniana
+ Plants much branched with most shoots fertile; petiole of middle stem leaves 0.8-5 cm; capitula usually with 20-60 disk florets (A. vestita with 12-20)   (27)
       
27 (26) Leaf blade 0.3-0.7 × 0.2-0.5 cm.   11 A. obtusiloba (see A. sect. Absinthium)
+ Leaf blade 2-10 × 0.7-8 cm   (28)
       
28 (27) Lobules of middle stem leaves entire, apex obtuse.   25 A. pontica
+ Lobules of middle stem leaves serrate or pectinate, apex acute   (29)
       
29 (28) Middle stem leaves abaxially yellow arachnoid pubescent, lobules linear or narrowly lanceolate; capitula with 40-60 disk florets.   29 A. stechmanniana
+ Middle stem leaves abaxially gray pubescent, often glabrescent, lobules serrate or pectinate; capitula with 12-40 disk florets   (30)
       
30 (29) Middle stem leaves triangular or elliptic-ovate in outline, lobules with triangular teeth; phyllaries puberulent or glabrescent; capitula with 20-40 disk florets.   28 A. gmelinii
+ Middle stem leaves ovate, elliptic-ovate, or suborbicular in outline, lobules deeply serrate; phyllaries densely pubescent; capitula with 12-20 disk florets.   30 A. vestita

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