All Floras      Advanced Search
FOC Vol. 2-3 Page 542, 572, 615, 730 Login | eFloras Home | Help
FOC | Family List | FOC Vol. 2-3 | Dryopteridaceae

18. Dryopteris sect. Dryopsis (Holttum & P. J. Edwards) Li Bing Zhang, Phytotaxa. 71: 18. 2012.

轴鳞鳞毛蕨组 zhou lin lin mao jue zu

Authors: Shiyong Dong & Maarten J. M. Christenhusz

Dryopsis Holttum & P. J. Edwards, Kew Bull. 41: 179. 1986.

Plants terrestrial, 25-150 cm. Rhizomes short, erect or rarely prostrate, apex covered by lanceolate scales. Fronds tufted; stipe stramineous, brown, or castaneous, scaly throughout; stipe scales brown or blackish brown, narrowly to ovate-lanceolate, never clathrate or iridescent (as those in Ctenitis). Lamina lanceolate to ovate, once pinnate with deeply lobed pinnae to amply 3-pinnate; basal pinnae oblong, lanceolate, or triangular, usually not or slightly longer than next pair, basal basiscopic lobe or pinnule mostly not enlarged; distal pinnae adnate to rachis and ± decurrent at their bases; veins free; lamina papery, adaxial surface with thick multicellular hairs; rachises and costae grooved and densely covered with ctenitoid hairs or sometimes with hair-scales (structure intermediate between hairs and scales) on adaxial surface, various scale types present or not on abaxial surface. Sori terminal or dorsal on veins, medial to submarginal, one line on either side of costules; indusia present, persistent; spores ellipsoid to spheroidal, perispore echinate or coarsely tuberculate. x = 41.

Sixteen species: tropical and subtropical Asia, southwest to S India and Sri Lanka, east to Japan and the Philippines, south to Malaysia and Indonesia, most diverse in the S and SE Himalaya; nine species (five endemic) in China.

1 Fronds 3-pinnate to 4-pinnatifid, with many free pinnules present; basal stipe scales broadly lanceolate or ovate-lanceolate, generally 2-5 mm wide; abaxial surface of costae with few scales except in D. ×holttumii; perispore coarsely tuberculate   (2)
+ Fronds 2-pinnate or rarely 3-pinnatifid, if present pinnules usually adnate to costae; scales on stipe base narrowly lanceolate, usually less than 1 mm wide; abaxial surface of costae copiously scaly; perispore sharply echinate   (4)
2 (1) Rachis sparsely scaly; indusia poorly developed and difficult to see, but sometimes fully developed and persistent or fugacious.   145 D. sphaeropteroides
+ Rachis densely scaly; indusia fully developed and persistent   (3)
3 (2) Sori throughout length of pinnules; abaxial surface of costae sparsely scaly.   143 D. maximowicziana
+ Sori confined to distal parts of pinnules; abaxial surface of costae copiously scaly.   140 D. ?olttumii
4 (1) Scales on abaxial surface of costae flat   (5)
+ Scales on abaxial surface of costae ?bullate   (6)
5 (4) Pinnules or pinna lobes with conspicuous hyaline margins.   138 D. clarkei
+ Pinnules or pinna lobes lacking hyaline margins.   144 D. transmorrisonense
6 (4) Scales on abaxial surface of costae slightly bullate; sori confined to distal parts of pinnules.   137 D. apiciflora
+ Scales on abaxial surface of costae obviously bullate; sori throughout length of pinnules   (7)
7 (6) Stipe stramineous or rarely dark brown; pinna lobes entire, crenate, or shallowly lobed, rarely pinnatifid; abaxial surfaces of lobe costae without or with a few scales.   142 D. kawakamii
+ Stipe castaneous and shiny; pinna lobes usually shallowly pinnate to deeply lobed, rarely entire; abaxial surfaces of lobe costae scaly   (8)
8 (7) Basal stipe scales few, ca. 0.5 cm; sori throughout length of pinnules.   141 D. heterolaena
+ Basal stipe scales copious, ca. 1 cm; sori usually confined to distal part of pinnules.   139 D. dulongensis

Lower Taxa


Flora of China @
Browse by
Advanced Search

Flora of China Home




 |  eFlora Home |  People Search  |  Help  |  ActKey  |  Hu Cards  |  Glossary  |