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FOC | Family List | FOC Vol. 2-3 | Dryopteridaceae

19. Dryopteris sect. Nothoperanema (Tagawa) Li Bing Zhang, Taxon. 61: 1206. 2012.

肉刺鳞毛蕨组 rou ci lin mao jue zu

Authors: Sugong Wu, Xiang Jianying, Li-Bing Zhang & David S. Barrington

Nothoperanema (Tagawa) Ching.

Plants terrestrial, medium-sized. Rhizome stout, erect or obliquely ascending. Fronds caespitose; stipe stout, with dense scales; scales lucid, ferruginous-brown or dark brown, ovate, lanceolate or subulate above, thick, entire, apex acuminate; lamina large, ovate-triangular, tripinnate or tripinnatifid; basal pinnae largest, subtriangular, base inequilateral; basiscopic pinnule largest, extending downward, pattern of pinnule arrangement such that basal pair anadromous; upper ones ± catadromous; from second pinnae lanceolate or broadly lanceolate, base equilateral; rachis with dense narrowly lanceolate or subulate scales; scales fuscous or nearly black, spreading; costa and costule glabrous on both surfaces, raised abaxially, grooved adaxially, with short, stout setae on each side of costae and at forking position of midribs; setae consisting of 3 rows of subquadrate stereids at base, 2 rows at middle, 1 row at apex. Venation free, same as Dryopteris type, veinlets not reaching margin, a hydathodal cell terminating veinlet. Lamina herbaceous or papery when dry, brown. Sori orbicular, terminating veinlets, 1 row on both sides of costa; indusia sometimes exindusiate, brown or dark brown, orbicular-reniform, firm, entire, attached by sinus, persistent or fugacious. Spores bilateral, reniform, surface verrucose or reticulate. n = 41.

About six species: C, E, and S Africa, E, NE, and S Asia, Himalaya, Indian Ocean islands (Mascarene Islands), Madagascar, New Guinea, and Pacific islands (including Hawaiian Islands); five species (two endemic) in China.

1 Rachis scales lanceolate and linear-lanceolate   (2)
+ Rachis scales long subulate   (3)
2 (1) Sori indusiate; rachis scales reddish brown; ultimate segments serrate apically.   149 D. hendersonii
+ Sori exindusiate; rachis scales dark brown; ultimate segments entire or undulate apically.   150 D. shikokiana
3 (1) Plants 120-150 cm tall; basalmost pinnae up to 38 cm; primary segments up to 17 cm; secondary segments up to 2 cm.   148 D. grandifrons
+ Plants 60-110 cm tall; basalmost pinnae 14-21 cm; primary segments 5-7 cm; secondary segments 1-1.6 cm   (4)
4 (3) Rachis scales nearly black when dried; pinnae acute apically.   146 D. squamiseta
+ Rachis scales brown to rust-colored when dried; pinnae acute or obtuse apically.   147 D. diacalpioides

Lower Taxa


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