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13. Polystichum sect. Macropolystichum Daigobo, Sci. Rep. Tokyo Kyoiku Daigaku. B 15: 61. 1972.

深绿耳蕨组 shen lü er jue zu

Polystichum sect. Prionolepia Daigobo.

Plants medium-sized to large, evergreen. Lamina often bipinnate, occasionally 1-pinnate to bipinnatifid, broadly lanceolate, ovate-lanceolate, or narrowly deltoid, thinly leathery or leathery, often with 1 or 2 proliferous bulbils toward rachis tip, occasionally with 1 proliferous bulbil toward apex tip of pinna rachis, some species without bulbils on rachis. Scales dimorphic to trimorphic, stipe often covered with dimorphic scales, base densely covered with large scales; large scales ovate, narrowly ovate or ovate-lanceolate, mixed with lanceolate to linear-lanceolate small scales, upper stipe and rachis covered with small scales, with few large scales; scales brown, thickly membranous, middle part often thickened and bright blackish brown; microscales light brown, small, subulate, basal part fimbriate, thinly membranous, often sparse on pinnae and pinnules abaxially. Sori often small, indusia small, easily falling off when mature.

About 17 species: tropical Asia and subtropical mountains, individual species widely distributed, north up to S Japan (Yakushima Island), some species west up to NW India, S Himalayan slopes; 14 species (six endemic) in China.

1 Rachis without proliferous bulbils   (2)
+ Rachis with proliferous bulbils   (7)
2 (1) Microscales lanceolate; sori abaxial on veinlets   (3)
+ Microscales filiform; sori terminal on veinlets   (4)
3 (2) Stipe scales up to 3 cm; pinnae 16-27 pairs.   73 P. mucronifolium
+ Stipe scales shorter than 2 cm; pinnae 8-17 pairs.   74 P. biaristatum
4 (2) Lamina acuminate and sterile at apex.   75 P. oreodoxa
+ Lamina cuspidate at apex   (5)
5 (4) Rachis with linear scales; lamina fertile at apex; pinnules oblong and falcate-oblong.   78 P. longipinnulum
+ Rachis with lanceolate scales at least at base of pinna rachis   (6)
6 (5) Lamina sterile at apex; pinnules deltoid-ovate or oblong.   76 P. semifertile
+ Lamina fertile at apex; pinnules narrowly deltoid-ovate.   77 P. fugongense
7 (1) Lamina 1-pinnate to bipinnatifid   (8)
+ Lamina bipinnate   (9)
8 (7) Lamina broadly lanceolate, nearly dimorphic, fertile lamina smaller; stipe scales with irregular teeth, teeth often forked or irregularly pinnately branching.   65 P. prionolepis
+ Lamina narrowly oblong-lanceolate, obviously monomorphic; stipe scales with regular simple teeth.   66 P. lentum
9 (7) Lamina 1-pinnatifid to pinnatisect   (10)
+ Lamina fully bipinnate   (12)
10 (9) Stipe scales brown; pinnules with microscales linear adaxially; rachis with 2 bulbils.   67 P. bigemmatum
+ Stipe scales mostly castaneous, at least castaneous at center; pinnules glabrous adaxially; rachis with 1(or 2) bulbils   (11)
11 (10) Pinnules and lobes of lower pinnae broadly rhombic-ovate or obovate, cuspidate aristate-spinulose.   68 P. chunii
+ Pinnules and lobes of lower pinnae ovate or oblong, obtuse-acuminate to long acuminate.   69 P. attenuatum
12 (9) Lamina nearly dimorphic, pinnae and pinnules of fertile fronds narrower and smaller; sori coenosori when mature, fully covering abaxial surface of pinnules.   70 P. pycnopterum
+ Lamina monomorphic; sori not coenosori when mature   (13)
13 (12) Pinnules shortly aristate-spinulose at apex and on distal margins.   71 P. disjunctum
+ Pinnules not aristate-spinulose, shallowly toothed, incised to repand on margin.   72 P. scariosum

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