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19. Polystichum sect. Sphaenopolystichum Ching ex W. M. Chu & Z. R. He, Fl. Reipubl. Popularis Sin. 5(2): 223. 2001.

细裂耳蕨组 xi lie er jue zu

Plants small to medium-sized, evergreen. Frond texture papery, thinly papery, or rarely thinly leathery. Stipe scales uniform, ovate to lanceolate, often appressed, rarely spreading. Lamina lanceolate, broadly lanceolate, ovate-lanceolate, or ovate, often bipinnatifid to bipinnatipartite, bipinnatisect, tripinnatifid, rarely tetrapinnatifid. Ultimate lobes often narrow, linear to spatulate, entire, with 1 veinlet per lobe. Sori 1 per lobe or ultimate lobe, small, abaxial or terminal on veinlets. Indusia falling off early or sori without indusia.

About 12 species: China, India, Myanmar, and Vietnam; 12 species (nine endemic) in China.

Polystichum sect. Sphaenopolystichum is recognized here based on morphology.

1 Scales on stipe and rachis open   (2)
+ Scales on stipe and rachis all or mostly appressed   (4)
2 (1) Lamina tripinnate to tetrapinnatifid; stipe scales dense, up to 10 ?5 mm, reddish brown.   144 P. wattii
+ Lamina bipinnatipartite to bipinnatisect, or lamina base bipinnate; stipe scales sparse, shorter than 2 mm, brown   (3)
3 (2) Lamina elliptic-lanceolate, green.   145 P. martinii
+ Lamina deltoid-lanceolate, dark green.   146 P. cuneatiforme
4 (1) Lamina tripinnate or tetrapinnate; ultimate lobes linear   (5)
+ Lamina 1-pinnate to bipinnate; ultimate lobes not linear   (6)
5 (4) Lamina narrowly elliptic or broadly elliptic-lanceolate, middle or upward broadest; pinnae approximate, up to 40 pairs, yellowish green, those below middle or above middle narrowed gradually toward lamina base; rachis scales narrowly lanceolate; pinna rachis stramineous or greenish stramineous; sori indusiate.   135 P. caruifolium
+ Lamina narrowly ovate, broadest below middle; pinnae often separate from each other, fewer than 25 pairs, dark green, those below middle not or only slightly narrowed toward lamina base; rachis scales broadly ovate; pinna rachis green; sori exindusiate.   136 P. alcicorne
6 (4) Lamina 1-pinnate; pinnae pinnatilobed to pinnatipartite   (7)
+ Lamina bipinnate; pinnules toothed or pinnatilobed to pinnatipartite   (9)
7 (6) Upper side of pinnae pinnatipartite to pinnatisect and often forming 1 or few free lobes.   141 P. auriculum
+ Upper side of pinnae pinnatifid only and without free lobes   (8)
8 (7) Pinnae 30-40 pairs, only a few basalmost pinnae slightly narrowed.   142 P. bifidum
+ Pinnae 15-23 pairs, those below middle of lamina gradually narrowed toward lamina base.   143 P. pseudolanceolatum
9 (6) Lamina elliptic-lanceolate, 5-9 cm at widest (at middle); pinnae asymmetrical at base; pinnules toothed or pinnatifid to pinnatipartite, narrowly cuneate, basal acroscopic ones largest and slightly auriculate   (10)
+ Lamina ovate or ovate-lanceolate, (7-)10-23 cm at widest (below middle); pinnae symmetrical or nearly symmetrical at base; pinnules pinnatilobed to pinnatifid, rarely pinnatipartite, broadly cuneate, basal acroscopic ones same size as others; ultimate lobes ovate-deltoid, rarely clavate-oblong   (11)
10 (9) Pinnules pinnatifid to pinnatipartite; ultimate lobes elliptic-clavate or spatulate.   137 P. christii
+ Pinnules only dentate-spinulose on margin, elliptic.   138 P. bissectum
11 (9) Pinnae often acute; lower part of pinna rachis often not winged; pinnules up to 2 cm, wider than 5 mm; sori medial or nearly medial.   139 P. tonkinense
+ Pinnae often acuminate or long acuminate; lower part of pinna rachis often narrowly winged; pinnules up to 1 cm, ca. 3 mm wide; sori submarginal.   140 P. excelsius

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