Philodendron daguense Linden & André, Ill. Hort. 18: 192, t. 79. (1871). Type: Colombia. Valle: Río Dagua, Wallis s.n.(holotype, K). Philodendron carderi hort. Gard. Chron. 10: 56. (1878). Type not seen. Philodendron pilatonense Engl., Bot. Jahrb. Syst. 37(11): 129. (1905). Type: Ecuador. Pichincha: Río Pilatón, May 1899, Sodiro s.n. (holotype, B; isotype, G, MO). Philodendron discolor K. Krause, Notizbl. Bot. Gard. Berlin-Dahlem 9: 273-274. (1925). Type: Peru. Junín: Prov. Jauja, in the valley of Río Masamerich (Río Pontachuela) confluent of Río Pangoa, above Rasthütte Calabaza, 1500–1600 m, 11°30S, 7 May 1913, Weberbauer 6663 (holotype, B; isotype, F).

Usually hemiepiphytic, rarely terrestrial or epiphytic; stem appressed-climbing, densely scaly or setose, trichomes green to greenish white; pre-adult internodes to 20 cm long, 1–2 cm diam.; adult internodes smooth, scurfy, minutely wrinkled, semiglossy to matte, 3 –10 cm long, 2–6 cm diam., about as long as broad or longer than broad, gray to brown, roots moderately numerous, mostly short, covered at least on one side with fine root hairs, drying dark brown, semiglossy, faintly appressed-scaly; cataphylls 10–30 cm long, unribbed, sometimes bluntly 1-ribbed, green or reddish, densely scaly or setose, persisting as a tangled network of fibers. Leaves erect-spreading; petioles 33–65(90) cm long, 3 cm diam. at base, 1.5 cm diam. at apex, subterete, purplish violet to brownish, surface densely scaly, the scales of two distinct types, short, broad, often lacerate scales interspersed with long acicular scales, the latter erect, or spreading then erect; blades broadly ovate-cordate, thinly coriaceous, moderately bicolorous, acuminate to narrow acuminate at apex (the acumen mostly inrolled, 1–2.5 mm long), cordate at base, 28–75 cm long, 19–60 cm wide (1–1.7 times longer than wide), (0.7–1.5 times longer than petiole), broadest below point of petiole attachment, upper surface dark green or bronze-green blackish green on new leaves), velvety to matte, drying dark brown, yellow-green or gray-green, lower surface pale green tinged red-purple between secondary veins (weakly glossy and purple-violet on new leaves), matte, drying yellow-brown to reddish brown; anterior lobe 9.6–51 cm long, 10–59 cm wide (1.9–3 times longer than posterior lobes); posterior lobes 8–21 cm long, 5–28 cm wide, broadly rounded to obtuse; sinus hippocrepiform to obovate or closed; midrib flat to sunken, slightly paler than surface above, convex, matte, darker than surface below; basal veins 6–8 per side, with (0)1–2 free to base, numbers 3–4 coalesced 1–5 cm; posterior rib not at all naked on small blades or naked for 0.5–6 cm; primary lateral veins 3–6(8) per side, departing midrib at a 50—55 degree angle, sunken, paler than surface above, round-raised to convex, darker than surface below; minor veins distinct and darker than surface below, arising from both the midrib and primary lateral veins; "cross-veins" conspicuous, in part raised below. Inflorescences 1–4 per axil; peduncles 5–25 cm long, 1–2 cm diam., medium green to purplish, usually 0.66–1.75 times longer than the spathe; spathe densely scaly or setose, 10–22 cm long, 4 cm diam.; spathe blade medium green, whitish or reddish green outside, white to pinkish inside; spathe tube reddish green, medium green, or dull purple-violet outside, red or pale reddish (dark green post-anthesis) inside; spadix 14.6 cm long; pistillate portion cylindrical to weakly clavate, 1.8–4.6 cm long, 1 cm diam. at apex, 8 mm wide at base; staminate portion 9.4 cm long; fertile staminate portion cream; pistils 2.6–3.3 mm long, 1.4–1.7 mm diam.; ovary 4–5-locular, 1.9–2.5 mm long, 1.4–1.7 mm diam., with axile placentation; locules 1.9–2.5 mm long, 0.5–0.7 mm diam.; ovule sac 1.8–2.2 mm long; ovules 20–24(34) per locule, arranged in series of 2, contained within translucent, gelatinous ovule sac, 0.1–0.2 mm long, as long as or longer than funicle; funicle to 0.2 mm long, adnate to lower part of partition, style 0.5–0.6 mm long, 1.4–1.7 mm diam., similar to style type B; style apex more or less flat; stigma more or less lobed, 1.4 mm diam., 0.2–0.3 mm high, covering entire style apex; the androecium truncate, prismatic, oblong, margins more or less 4–6-sided, 0.8–0.9 mm long, 2–2.5 mm diam. at apex; thecae oblong, 0.3–0.4 mm wide, more or less parallel to one another; sterile staminate flowers blunt, irregularly 4–5-sided, sometimes clavate or prismatic, 1.6–1.8 mm long, 1.4–1.7 and 1.9–3.4 mm wide. Infructescence with ripe berries white.

Lectotype: Locality unknown. Schott Drawings 2757--2759 (microfiche 53:a8﷓10). [Type designated here]. Philodendron daguense Linden & André, Ill. Hort. 18: 192, t. 79. (1871). Type: Colombia. Valle: Río Dagua, Wallis s.n.(holotype, K). Philodendron carderi hort. Gard. Chron. 10: 56. (1878). Type not seen. Philodendron pilatonense Engl., Bot. Jahrb. Syst. 37(11): 129. (1905). Type: Ecuador. Pichincha: Río Pilatón, May 1899, Sodiro s.n. (holotype, B; isotype, G, MO). Philodendron discolor K. Krause, Notizbl. Bot. Gard. Berlin-Dahlem 9: 273﷓274. (1925). Type: Peru. Junín: Prov. Jauja, in the valley of Río Masamerich (Río Pontachuela) confluent of Río Pangoa, above Rasthütte Calabaza, 1500--1600 m, 11°30S, 7 May 1913, Weberbauer 6663 (holotype, B; isotype, F).

Esmeraldas: 6 km beyond bridge over Río Esmeraldas near San Mateo, road to Esmeraldas airport, Río Mutile, 80 m, 0°52'N, 79°33'W, Croat 55609 (MO); Zapallo Grande along Río Cayapa, 200 m, 0°48'N, 78°54'W, Barfod et al. 48191 (AAU, MO); Awá Reserve, Community of Mataje 200 m, 1°15'N, 78°40'W, Beck 1773 (MO); Bilsa Biological Reserve, Montañas de Mache, 35 km W of Quinindé, 5km W of Santa Isabela, 400-600 m, 0°21'N, 79°44'W, Pitman & Bass 914 (MEXU, MO, QCNE), Pitman & Bass 1031 (MO, QCNE); Lita-San Lorenzo Road, 14.2 km W of Río Lita Bridge (below Lita), 425 m, 0°52'11"N, 78°27'16"W, Croat et al. 82360 (MO). Guayas: Junction of Provinces Guayas, Cañar, Chimborazo, and Bolívar, Near the village of Bucay, 305-380 m, Camp E-3740 (NY). Los Ríos: Centinela, 0°36'S, 79°18'W, Iltis E-80, E-80A (WIS); Hcda. Clementina, cerro Samana, 1°40'S, 79°21'W, Cornejo et al. 4974 (GUAY, MO ); Hac. Clementina, 750 m, Harling 488 (S). El Oro: Piñas-Santa Rosa above El Placer, 400-700 m, Harling et al. 15523 (GB); 10.5 km W of Piñas on road to Arenillas, 975 m, Thompson 152 (MO); road from Arenillas to Piñas, 502 m, 3°40'S, 79°45'W, Thompson 370 (MO). Carchi: Parroquia Tobar Donoso, sector Sabalera, Reserva Indigena Awá, 650-100 m, 1°0'N, 78°24'W, Tipaz et al. 1466 (MO, QCNE); Reserva Etnica, Awá, Parroquia Chical. Centro Gualpi Medio, 900 m, 1°2'N, 78°16'W, Aulestia & Grijalva 1151 (MO, QCNE); Chical, 1200-1250 m, 0°56'N, 78°11'W, Thompson & Rawlins 1002 (CM, MO). Imbabura: in valley of Río Mira on border with Carchi Province, between Ibarra and Lita, 2.5 km E of Lita, 750-775 m, Croat 38975 (MO). Pichincha: Reserva Maquipucuna, Hacienda El Carmen, trail up Gregoire's Hill, ca 6 km airline SE of Nanegal, 1500 m, 0°7.5'N, 78°38'W, Webster & Delprete 27177 (MO); 11 km W of Tandapi, trail along Chictoa River, tributary of Río Pilatón, 1350-1550 m, Gentry et al. 12119 (S); Reserva ENDESA, 9 km N of Km. 113 on Quito- Puerto Quito Hwy, 750 m, 0°5'N, 79°2'W, Croat & Rodríguez 61488 (MO); Pacto-Nuevo Azuay, 10 km N of La Esperanza, 1150 m, 0°12'N, 78°46'W, Holm-Nielsen et al. 24569 (AAU, MO); El Paraiso-Saguangal, 3 km from El Paraiso, 1500 m, 0°19'N, 78°46'W, Øllgaard et al. 37803, 37835 (AAU); 1500 m, 0°10'N, 78°46W, Øllgaard 37777 (AAU); Km 113, Nanegalito-Puerto Quito Road, then north on road to ENDESA Reserve, 602 m, 0°4.52'N, 79°1.44'W, Croat 82788 (MO); Río Pilaton, Sodiro s.n. (MO sheet 1763728 and -29) (MO). Reserva Maquipucuna, El Pacchal, S slopes of Cerro Campana, ca 4-5 km airline SE of Nanegal, 1300-1500 m, 0°8'N, 78°38'W, Webster & Kelch 27949 (DAV, MO); along road between Nono and Nanegal, NW of Quito, 8 km NW of Nanegalito, 1280 m, Croat 38886 (MO). Cotopaxi: Road between Quevedo & El Corazón, 63.4 km SE of Quevedo; 6 km NW of El Corozon; above Río Angamarca;1030 m, 1°7'S, 79°7'W, Croat 55769 (MO); Tenefuerste, Río Pilalo, Km 52-53 between Quevedo and Latacunga, 750-900 m, Dodson & Dodson 12932 (MO, SEL). Azuay: Río Putucay-Río Norcay, between Chacanceo and Río Blanco, on road to Molleturo, 1220 m, Steyermark 52811 (NY); Sanagüín, Hacienda Yubay-Hacienda San José de Caimotán, along Río Patul, 850 m, Steyermark 52744 (NY). Sucumbios: along road between Baeza and Nuevo Loja (Lago Agrio), along slope above road and Río Aguarico, 67 km E of Baeza, 2.7 km W of Río Azuela, 1270 m, 0°10'S, 77°41'W, Croat 72490 (CM, MO, SAR). Napo: Proyecto Hidroeléctrico Coca, Punto Sta.4, Márgen derecha del Río Quijos, ca 10 km al sur de Reventador, 1450 m, 0°8'S, 77°30'W, Palacios 5984 (MO, QCNE); Cantón El Chaco. Márgen derecha del Río Quijos, Finca "La Ave Brava" de Segundo Pacheco. 1800-1900 m, 0°12'S, 77°39'W, Palacios 5338 (MO); new road to Loreto (56 km S of Baeza); 18 km E of junction with Baeza-Tena road, 1200 m, 0°50'S, 77°41'W, Hammel & Wilder 17273 (DUKE,ENCB,MO); San Francisco Borja, Croat 67418 (MO); along road between Baeza and Lago Agrio at 125.3 km W of Lago Agrio, 1500 m, Croat 49468 (MO); along road between Baeza and Lago Agrio, 107 km W of Lago Agrio, 1470 m, Croat 49490 (MO); 2.5 km NNE of Río Reventador on Baeza-Lago Agrio Rd, 1660 m, MacBryde 1469 (MO); Cosanga, 1900 m, Boeke & McElroy 423 (MO); Baeza-Tena Rd., southern slope of Cordillera de Guacamayos above Jondachi, 1700 m, Harling & Andersson 16353 (GB). Pastaza: Shell-Mera, 5.3 km NW of Center of Shell, along gravel road 1.1 km N of hwy, 1180 m, 1°27'S, 78°4'W, Croat 73480 (MO); Mera, 1160 m, Harling 3141 (S); El Japo, 1250 m, Harling 3422 (S). Morona-Santiago: Cordillera del Cóndor, Cuangos, 20 km east of Gualaquiza, near disputed Peru-Ecuador border, 1480 m, 3°29'S, 78°14'W, Gentry 80126 (MO, QCNE); above Ganazhuma, 1825- 2200 m, 3°27'S, 78°50'W, Thompson 341 (MO); 9-10 km SE of San Juan Bosco, 1540-1600 m, Gentry et al. 30897 (MO, SEL). Zamora-Chinchipe: along road between Loja and Zamora, at Río Zamora, 1610 m, 4°5'S, 79°0'W, Croat 72694 (MO).