Herbs, perennial, from creeping rhizomes, aerial stems usually proximally tomentose and distally stipitate glandular. Leaves simple, alternate, basal and cauline, petiolate (proximal) or sessile (distal); blades ovate to triangular or suborbicular, abaxially tomentose, adaxially glabrate, bases mostly truncate to cordate or hastate, margins coarsely dentate or lobulate to denticulate or entire. Synflorescences of lax panicles. Capitula disciform; involucre campanulate; phyllaries 5-7, 1- or 2-seriate, subequal, ovate to oblong, herbaceous, apices obtuse; receptacles convex, smooth, glabrous, epaleate. Marginal florets female, corollas white or ochroleucous, unequally and deeply 5-lobed; disk florets functionally male, corollas white or ochroleucous, 5-lobed, actinomorphic, corolla tube longer; anther basal appendages (relatively short) entire, apical appendages narrowly triangular; style undivided, style branches papillate outside. Achenes clavate to obovoid, not beaked, obscurely ribbed, covered with stout red stipitate glands, glands densest in upper half, achenes of inner florets usually glabrous; carpopodium annular; pappus absent. 2n = 42, 46, 92.

About five species: E Asia, Central, North, and South America; one species in China.

(Authors: Chen Yousheng (陈又生); D. J. Nicholas Hind)