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Juglandaceae A. Richard ex Kunth


Description from Flora of China

Trees or rarely shrubs, deciduous, semievergreen, or evergreen, monoecious or rarely dioecious; bark tight (or exfoliating). Branchlets with solid or chambered pith. Terminal buds subglobose or ovoid to oblong, naked or with scales. Stipules absent. Leaves alternate (or opposite), odd- or even-pinnate, sometimes trifoliolate, rarely simple; leaflets with glandular, peltate scales, often resinous and aromatic, particularly conspicuous abaxially on young leaves and twigs, margin serrate or rarely entire. Inflorescences pendulous or sometimes erect, lateral or terminal, on reduced shoots arising on branchlets of previous year (old growth) or on current year’s growth (new growth), of several types: androgynous panicle with male, lateral spikes and female, central spike; androgynous panicle with male, mainly lateral spikes and female, central spike male at apex; cluster of male spikes and solitary female spike; or solitary male and female spikes. Flowers unisexual, anemophilous, rarely entomophilous. Male flowers subtended by an entire or 3-lobed bract; bracteoles 2 or absent; sepals 0-4, adnate to receptacle when present; stamens 3-40(-100), inserted on receptacle; filaments short to nearly absent, free or united at base; anthers glabrous or pubescent, 2-loculed, dehiscing longitudinally. Female flowers with an entire or 3-lobed bracts; bracteoles 2 or 3 (or absent); sepals 0-4, adnate to ovary, free at apex; gynoecium of 2 carpels united into an inferior ovary, 1-loculed, but at base 2-4(-8)-loculed; style 1, short or elongate, rarely absent; stigmas 2, carinal or commissural, sometimes 4-lobed, plumose or fleshy; ovule 1, orthotropous. Fruiting spike elongate, and pendulous or short and erect, rarely conelike. Fruit a drupelike nut, 2-4(-8)-chambered at base, with a dehiscent or indehiscent husk, or a 2- or 3-winged or disc-winged nutlet. Seed solitary, without endosperm. Cotyledons 4-lobed, much contorted. Germination hypogeal or epigeal. 2n = (28), 32, (64).

Nine genera and 60 or more species: mostly in temperate and subtropical regions of the N hemisphere; seven genera (one endemic) and 20 species (seven endemic, one introduced) in China.

Kuang Ko-zen & Lu An-ming. 1979. Juglandaceae. In: Kuang Ko-zen & Li Pei-chun, eds., Fl. Reipubl. Popularis Sin. 21: 6-44.

(Authors: Lu Anmin (路安民 Lu An-ming) ; Donald E. Stone , L. J. Grauke)

Lower Taxa


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