Herbs perennial, erect, pubescent or villous. Basal leaves long petiolate; stem leaves alternate, entire. Cymes ebracteate. Calyx 5-parted to base; lobes lanceolate to linear-oblong, slightly enlarged in fruit. Corolla funnelform; tube usually longer than calyx; throat appendages elongated, curved, or oblong, rarely reduced, becoming ovate, entire at apex; lobes of limb subvertical or spreading, obtuse. Stamens inserted below throat; anthers elongated, frequently hastate at base, exserted from throat. Style filiform, exserted, thickened and persistent in fruit. Gynobase short conical. Nutlets dorsiventrally compressed, ovate, ca. 6 mm, abaxially discoid with glochids; attachment scar above middle adaxially, ovate, firmly coherent to gynobase.

About ten species: C and W Asia; one species in China.