Plants large, terrestrial. Rhizome long and creeping, aerial shoots woody and vinelike, terete, with sparse leaves. Sterile branches soft, yellowish green, terete, dichotomously branched with well-differentiated lateral branchlets much branched; leaves spirally arranged, but leaf base twisted and making branchlets complanate, dense, ascending, subulate, herbaceous. Fertile branches soft, reddish brown, branchlets flattened, dichotomously branched with well-differentiated lateral branchlets; leaves spirally arranged, sparse, appressed, scalelike; bracts similar to leaves on aerial shoots but smaller; strobili 6-26 per peduncle and terminal on multi-dichotomously branched peduncles forming a cone shape, with erect peduncles and pedicels, bent, reddish brown; sporophylls broadly ovate, imbricate, thickly membranous, with irregular teeth on margin, apex acute, with membranous long awn. Sporangia borne in axils of sporophylls, enclosed, yellow, reniform.

One species: widely distributed in tropics and subtropics of Asia extending eastward to Papua New Guinea.